Orthogonius euthyphallus bolavenensis Tian & Deuve

Tian, Mingyi & Deuve, Thierry, 2013, Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini), ZooKeys 349, pp. 81-100 : 88-89

Orthogonius euthyphallus bolavenensis Tian & Deuve ssp. n. Figs 1, 6-7, 16-17

Description.

Body stout, strongly shiny. Habitus as in Figs 6-7.

Length: 16.5 mm; width: 6.5 mm.

Black, antennomeres 4-11, mandibles, palps and tarsomeres dark brown to brown.

Macrosculpture: Head densely punctured in both sexes, elytra densely punctured in male, (rather sparser in female); pronotum much less punctured in male, without puncture in female; head obscure striate on frons, smooth on vertex; pronotum faintly striate or not; underside surface smooth and glabrous, impunctate.

Microsculpture: Engraved meshes densely isodiametric on elytra, faint on head and pronotum.

Head longer than wide, HL/HW=1.17-1.18, eyes large and prominent, frons and vertex convex, frontal impressions large and deep, extending on clypeus; clypeus bisetose, basal portion deeply furrowed; labrum slightly broad at apical margin, sexsetose; palps rather stout, subcylindrical, maxillary palpomere 3 as long as 4 which is glabrous, palpomere 3 with two short setae at apex; labial palpomere 2 longer than 3, bisetose on inner margin, with several additional short setae at subapex and apex, palpomere 3 with a few tiny setae; ligula small, bisetose at apex; mentum without tooth, each of mentum and submentum bisetose, palpiger asetose. Antennae rather stout, not extending over the middle of elytra; pubescent from basal 1/4 of antennomere 4; antennomere 3 almost as long as 4, and 1.55 times longer than 2; antennomere 1 unisetose on subapex.

Pronotum rather broad though transverse, PW/PL=1.49-1.52, disc moderately convex, fore and basal margins well beaded, sides evenly expanded, widest at about middle; lateral expanded margin wide and hardly reflexed, better defined in female than in male; both transversal impressions indistinct, basal foveae shallow, median line clear.

Elytra elongate ovate, EL/EW=1.57-1.60, widest at about middle, where sides slightly expanded, base completely bordered, apex broadly sinuate, inner angle nearly rectangular, faintly denticulate; striae deep, intervals convex, subequal in width in middle portion, interval 3 with basal and middle setiferous pores, the subapical one absent.

Legs slender, fore tibia distinctly expanded at apex, apex obliquely truncate, outer margin smooth, not subserrate; middle and hind coxae smooth and glabrous in median portion; middle tibia slender and slightly expanded at apex, strongly curved in male, while normal in female; hind femur moderately expanded, with two setae posteriorly; hind tibia slightly expanded at apex, apical spurs long and sharp; fore tarsi distinctly wider than middle and hind ones, middle wider than the hind; hind tarsomeres 1 and 3 much longer than 2 and 4 respectively, length ratio of tarsomeres 1-4 as 2.57, 1.86, 1.28 and 1.0; tarsomere 4 asymmetrically emarginat, with longer lobe as deep as 1/3 of the joint; all tarsal claws strongly pectinate.

Prosternal process well bordered at apex, abdominal ventrite VII of male narrowly and deeply emarginate at apical margin.

Male genitalia (Figs 16-17): Slender and quite straight, expanded medially, gently sinuate on ventral margin, apex stout and blunt in profile; in dorsal view, broad at subapical portion, the apical lamella slightly wider than long, broad at tip.

Remarks.

Similar to the nominate subspecies, but darker and broader (in particular pronotum and elytra), and the apical lamella is more slender in dorsal view.

Material examined.

Holotype: male, "Coll. I. R. Sc.N.B/ purchased from Mr. B. Makovsky, I.G. 31.969" and "Laos South, Bolaven Plateau, Ban Itou, 23.-27. V. 2007, B. Makovsky Lat.", in IRSNB.

Paratype. 1 female, idem, in IRSNB.

Etymology.

Refers to its type locality.

Distribution.

Southern Laos. Known only from the type locality (Fig. 1).