Osmylidia glastrai, Makarkin & Archibald & Mathewes, 2021

Makarkin, Vladimir N., Archibald, S. Bruce & Mathewes, Rolf W., 2021, New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks, Zootaxa 4980 (1), pp. 142-156 : 144-145

Osmylidia glastrai sp. nov.

Fig. 1

Type material. Holotype SR 02-22-15 (part) collected by Joshua Glastra, August 24, 2002; deposited in SR. A wellpreserved almost complete forewing.

Type locality and horizon. Exposure B 4131 of the Tom Thumb Member of the Klondike Mountain Formation in the town of Republic, Washington, U.S.A.; Eocene (late Ypresian) .

Etymology. From the surname of Joshua Glastra, collector of this fossil.

Diagnosis. Forewings may be distinguished from those of other species of the genus by faint colour patterning ( O. picta : strongly spotted; O. requieta : without spots).

Description. Forewing ca. 17 mm long, ca. 5.7 mm wide. Costal space broad, with subcostal veinlets simple, closely spaced. Sc fused with RA far from wing apex. Sc+RA veinlets simple, closely spaced. Subcostal space narrow; crossveins not detected. RA space relatively narrow, with five preserved crossveins, rather regularly spaced; one distal crossvein probably not preserved. RP originates rather close to wing base, with 14 pectinate branches, one of these (RP10) deeply forked. M forked distad origin of RP1, proximad origin of RP2. All crossveins (beside one proximal) in radial and radio-medial spaces arranged in three gradate series: proximal series with five crossveins; intermediate series with eight crossveins; outer series with seven preserved crossveins (partly fragmentarily preserved). Basal crossvein in radio-medial space connects R, M. Both MA, MP pectinately branched distally, with three branches. Four crossveins between MA, MP; five preserved crossveins between MP, CuA. CuA pectinately branched, with three branches. CuP pectinately branched, with five preserved branches (probably six or seven in life). One preserved crossvein detected between CuA, CuP. One distal crossvein between CuP, A1. A1 relatively long, with five pectinate, simple branches. A2 relatively long, with four preserved pectinate, simple branches. Wing colouration: irregular spots throughout wing (surrounding some crossveins, in subcostal space and along margin), most prominently three between MP, CuA, one along margin near termination of CuP, A1, and two on pterostigma.

Remarks. The species has a five-branched CuP and, therefore, is assigned to Osmylidia (see genus diagnosis) but only tentatively, as this character is unknown in one fossil genus, and Osmylidia is barely distinguished from extant genera solely by forewing venation.

Figures

FIGURE 1. Osmylidia glastrai sp. nov., holotype forewing SR 02-22-15. A, photograph; B, drawing. Scale bar = 5 mm.