Paleogenia waichertae Loktionov, Lelej et Perkovsky, 2023

Loktionov, Valery M., Lelej, Arkady S., Vasilenko, Dmitry V. & Perkovsky, Evgeny E., 2023, Discovery of a new extinct spider wasp (Hymenoptera, Pompilidae) from Eocene Rovno amber, Zootaxa 5352 (3), pp. 426-432 : 427-430

Paleogenia waichertae Loktionov, Lelej et Perkovsky , sp. nov.

Figs 1–10

Type material. Holotype, ♁: SIZK K-3286, Klesov, Rovno amber, late Eocene. Syninclusions: SIZK K-3287: Diptera (female of Ceratopogonidae ), SIZK K-3288: Coleoptera, SIZK K-3289: Archaeognatha ( Machilidae) , 2 Hymenoptera (1 Mymarommatidae ), 5 Acari (larvae of Parasitengona ) . Paratype, ♁: SIZK K-8178, Klesov, Rovno amber, late Eocene. Syninclusion: SIZK K-8179: Diptera .

Diagnosis. Male of the new species clearly differs from those of all other congeners in the combination of the following characters: the tibial spur 0.8–0.9 × as long as the first tarsomere; the apical portion of the hypopygium well-exposed beyond the tergum 7 and the sternum 6, rod-shaped and slightly widened apically, bearing erect setae in ventral view ( Figs 5 , 6 ); the gena 0.2 × wider than the eye in lateral view ( Fig. 8 ). The female is unknown.

Description. MALE. Body length: 3.2 mm (holotype), 3.0 mm (paratype). Head without noticeable erect setae. Eyes large; eye 0.48 × as wide as median interocular distance (head in frontal view) ( Fig. 6 ). Clypeus large, convex in dorsal view, trapezoid; 0.58 × as long as wide; its apical margin nearly straight; its posterolateral corners rounded ( Fig. 6 ). Mandible apex bidentate. Gena narrow, 0.2 × as wide as eye (head in lateral view) ( Fig. 8 ). Antenna shortened, with 11 cylindrical flagellomeres ( Figs 2 , 6, 8 ); apical flagellomere pointed.

Metapostnotum well developed, long ( Figs 8, 9 ). Propodeum smooth, with dense micropubescence, in lateral view gently sloping, its posterior surface nearly flat ( Figs 3 , 9 ).

Tergum 1 not petiolate basally, gradually widening towards apex ( Fig. 3 ). Terga with dense micropubescence ( Fig. 3 ). Apical portion of hypopygium well exposed beoynd tergum 7 and sternum 6, rod-shaped and slightly widened apically, bearing erect setae in ventral view ( Figs 5 , 6 ).

Legs without noticeable erect spines ( Figs 4 , 6, 10 ). Tibia with short apical spine-like setae. Tibial spur of meso-and metalegs 0.8–0.9 × as long as basal tarsomere ( Figs 4 , 10 ). Tarsal claws very small, inner tooth indistinct.

Wings crumpled (characters are invisible).

FEMALE is unknown.

Distribution and age. Rovno amber, Ukraine. Priabonian, 33.9-37.8 Ma, Upper Eocene ( Colombo et al. 2021).

Etymology. The new species is named for Cecilia Waichert (Complexo Biopráticas, Universidade Vila Velha, Vila Velha, Brazil), one of the most active pompilid researchers that made a great contribution to the taxonomy of extant and extinct taxa.

Remark. Size of the piece and syninclusions clearly indicates that holotype was caught in the resin trap at the base of the trunk (see Perkovsky et al. 2012).

Figures

FIGURES 1–5. Holotype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-3286. 1, habitus, total dorsolateral view. 2, head and antenna, frontolateral view. 3, propodeum and metasoma, dorso-lateral view. 4, meso- and metatibia and tarsi, lateral view (arrow points hypopygium). 5, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale).

FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale).