Pampasatyrus quies (Berg, 1877) (Satyrus)

Matz, Jess & Brower, Andrew V. Z., 2016, The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes, Zootaxa 4125 (1), pp. 1-108 : 58-59

Pampasatyrus quies (Berg, 1877) (Satyrus)

( Figs. 9 F; 19F–H; 28)

Lectotype: (male) MNCN (photo examined) Type location: Carmen de Patagonia, Buenos Aires Province, Argentina = Cosmosatyrus quies f. plana Weymer, 1911 Holotype: unknown

Type location: Uruguay

Distribution. Found in eastern Paraná State, Brazil, southwest through Uruguay to the southernmost part of Buenos Aires Province, Argentina at nearly sea level to 3000m, from December to March and possibly as late as May in the southernmost part of its range ( Fig. 28 ).

Diagnosis. Most similar to Pampasatyrus gyrtone , but slightly smaller, with the pupils of the ocelli on the forewing smaller and white rather than blue, and with the median border of the hindwing postmedian band scalloped and white rather than nearly straight and yellow. Projections on the aedeagus in P. qu i e s protrude from a flange that extends laterally in a gentle arc on both sides and are more wing-like in P. gyrtone . Valvae closely resemble that of P. glaucope , but are rounder at the distal end.

Redescription. Head: Antennae 8–9mm with cream and chocolate scales, terminating in a spatulate club. Eyes oval and naked, length approximately 1.2 times width. Male palps white at the base to chocolate at the distal end of the dorsal side with a longitudinal white stripe along the median and chocolate with chocolate and cream piliform scales on the ventral side. Female palps almost completely cream to white with the distal end of the dorsal side chocolate and chocolate to dark chocolate piliform scales on the ventral side. Terminal palp segment cylindrical and a little more than one-third the length of the second segment.

Thorax bearing iridescent black and chocolate brown scales with chocolate brown piliform scales in the males, the females similar with the addition of cream scales and cream piliform scales. Forelegs extremely reduced with the tarsi clublike and unsegmented in both sexes. Midlegs and hindlegs with four rows of amber spines on the tibia and tarsus.

Forewing ( Fig. 9 F): Wingspan 30–32mm. Termen nearly straight to slightly convex and the distal end of the discal cell widely U-shaped. Males with no visible androconial patch. Dorsal side tan to coppery brown with fringe scales in chocolate brown with a ripple pattern in dark chocolate striations along the costa and apex. Discal cell may have a slight peach blush that extends into the postmedian band. Postmedian band tawny to tan with nearly straight to barely crenate edges in tan to chocolate brown. Ocellus between M1-M3 black, round, bipupillate, and ringed in daffodil yellow. Another black, round ocellus ringed in daffodil yellow appears between CuA1-CuA2 and may be unipupillate. Ventral side chocolate and may fade to tawny toward the costal edge in some specimens. A ripple pattern appears in dark chocolate striations over most of the wing, but more strongly along the costal and subterminal edges. Apex and subterminal band are highlighted in white and a peach-brown blush may appear over the cubital edge of the discal cell and the cubital veins. Postmedian band tan to tawny or chocolate with both median and subterminal border dark chocolate brown and nearly straight. Median edge of the postmedian band daffodil yellow between M1-CuA2. Apical ocellus between M1-M3 similar to the dorsal side, but bolder. A small, round, black ocellus that shares the daffodil yellow ring with the M1-M3 ocellus may appear between M3-CuA1as a distinct ocellus or as fused to the M1-M3 ocellus. An ocellus between CuA1-CuA2 appears as in the dorsal side, but bolder and unipupillate.

Hindwing ( Fig. 9 F): Wing trapezoidal, termen slightly convex, barely scalloped, almost entire, and the inner margin slightly excavated between the anal vein and 1A+2A. Dorsal side and fringe scales similar in color to the forewing with the postmedian band appearing as a red-orange to tawny patch with a chocolate to dark chocolate deckle-edged subterminal border. Ventral side chocolate to taupe with dark chocolate striations in a ripple pattern over the entire wing. Postmedian band cream to tan with dark chocolate borders, the median border scalloped with the space between scallops at M3 extending a little farther toward the discal cell into a white triangle and the subterminal border, when distinguishable, deckle-edged and thicker than the median border. Black lenticular ocelli appear between Rs-M1 and M1-M2, the latter larger than the former and both ringed in daffodil yellow. The ocellus between M1-M 2 may be unipupillate. Other, similar ocelli may appear between M2-M3, CuA1-CuA2, and rarely between M3-CuA1. Veins may be highlighted in tan to white.

Male genitalia ( Fig. 19 F–H): Uncus widest where it joins the tegumen, narrowing gradually to a blunt distal end, and approximately 1.3X the length of the tegumen. Gnathos acute and about three-eighths the length of the uncus. Valvae generally wide, the proximal half acute triangular with a blunt proximal end. Distal half with the ventral side edge nearly straight and the dorsal side edge proceeding from the distal end in a wide curve that begins at about a 60° angle from the ventral side edge and flattens as it approaches the median. Aedeagus widest at the distal one-third bearing dentate projections on each side, slightly narrower at the distal end, and narrowing toward the proximal end to about half the width at the proximal one-third to a truncate terminus.

Specimens examined. Uruguay, Montevideo Province, (CUIC) 1 male; Argentina, Rio Negro Province, (MTSU) JMC0809

Figures

FIGURE 9. Composite habitus images of Pampasatyrus species. The left side of each specimen is the dorsal surface, the right side is the ventral surface. A: Pampasatyrus gyrtone (Berg, 1877) male (BMNH # 809453); B: Pampasatyrus edmondsii (Butler, 1881) n. comb. male (MZUJ); C: Pampasatyrus glaucope (C. Felder & R. Felder, 1867) male (BMNH # 789978); D: Pampasatryrus gorkyi Pyrcz, Cerdeña and Zacca, 2014 holotype male (scale bar = 1 cm; reproduced from Cerdeña et al., 2014, open access); E: Pampasatyrus nilesi (C. Felder & R. Felder, 1867) male (MZUJ); F: Pampasatyrus quies (Berg, 1877) male (MZUJ).

FIGURE 19. Male genitalia of Pampasatyrus species. See Fig. 2 for labels of parts. All drawings to the same scale. A, B: Pampasatryrus gorkyi Pyrcz, Cerdeña and Zacca, 2014. Photos of holotype male (MUSM), with lateral and dorsal views of aedeagus (B) (modified from Cerdeña et al., 2014, open access); C – E: Pampasatyrus nilesi (C. Felder & R. Felder, 1867) (MGCL), (D) showing ornamentation on the aedeagus; F – H: Pampasatyrus quies (Berg, 1877) (BMNH # 808376), showing ornamentation on the aedeagus (G).

FIGURE 28. Distribution of locality records for Pampasatyrus species. Note that the locality for P. glaucope is overlain by a record for P. reticulata.