Persicaria omoloica Doweld, 2017

Doweld, Alexander B., 2017, New names in Fallopia, Persicarioipollis, Polygonum and Reynoutria (Polygonaceae), living and fossil, Phytotaxa 308 (1), pp. 66-79 : 70-71

8. Persicaria omoloica Doweld , sp. nov.

Type:— RUSSIAN FEDERATION. Irkutsk region, Ust’- Kutsk district, Khapchan-khaja , left bank of Omoloj river (holotype BIN 19 /1- K550 /1! ( LE); figured by Arbuzova 2006: pl. 92, fig. 11) .

–“ Polygonum omoloicum ” in Dorofeev (1972: 94), nom. inval. (Art. 40.1).

–“ Persicaria omoloica ” in Arbuzova (2006: 16), nom. inval. (Art. 43.1).

Diagnosis:— Fruits 2.0–2.35 × 1.9–2.0 mm, ellipsoidal, sometimes apically widened, convex-planate or biconvex, initially bloated. Apex rounded, with a small remain of stylus; base shortly wedge-shaped, acicular, accrescent by perianth remains; side slightly tight. Fruits black, smooth; fruit surface foveolate in longitudinal series, forming slightly differentiated grooves. Pericarp 0.11–0.13 mm thick, 2-layered; exocarpium 1-layered, lignified, consisting of large, radially elongate cells, outer and inner cell walls thin, radial cell walls thick, porate, highly thickened in the upper 1/3 fruit part; mesocarpium thin, consisting of tangentially elongate and flattened cells.

Eponymy:— From the name of Omoloj river in Siberia, locus classicus.

Stratigraphy:— Miocene.

Status:— Fruits.

IFPNI registration lsid:— E0CF22F9-98D3-4999-A523-58BD9C06BC12

Note:— Dorofeev (1972: 94) missed the indication of the holotype, so, “ Polygonum omoloicum ” was not validly published (Art. 40.1) (see IFPNI: http://fossilplants.info/names/ 4AA72473-4010-4B97-8796-F2D606902BF3). Arbuzova (2006: 16–17) found only one fossil specimen in the preserved Dorofeev’s palaeocarpological collections at LE, but she erroneously thought that if the single figured specimen preserved in author’s collections, it might serve as a holotype. In fact, Art. 40.1 explicitly requires that the indication of the holotype must be given in the original publication. Arbuzova (l.c.) invalidly transferred the Dorofeev’s name into Persicaria , but failed to provide a required diagnosis in English or Latin for fossils (Art. 43.1). As a result, conditions for the valid publication of a new fossil-species were not fulfilled, and the combination, which in fact was an inadvertent publication of a new fossil-species, was not validly published. Hence, a new fossil-species is here described as Persicaria omoloica .