Philodromus tuvinensis, Szita & Logunov & O, 2008

Szita, É., Logunov, D. & O, P., 2008, A Review Of The Histrio Group Of The Spider Genus Philodromus Walckenaer, 1826 (Araneae, Philodromidae) Of The Eastern Palaearctic Region, Acta Zoologica Academiae Scientiarum Hungaricae 54 (1), pp. 23-73 : 46-50

publication ID	https://doi.org/ 10.5281/zenodo.12585274
persistent identifier	https://treatment.plazi.org/id/03DD87CC-E51A-D66D-7408-FA39FFE0F9E1
treatment provided by	Felipe
scientific name	Philodromus tuvinensis
status	sp. nov.

Treatment

Philodromus tuvinensis View in CoL sp. n.

(Figs 27–29, 41–42, 73–74, Map 3)

Type material: The m holotype ( ZMMU) from Russia, Tuva, Erzin Distr., SE shore of Tere-Khol’ [=Tore-Khol’] Lake , Eder-Elezin Sands , Sharaa Stand , poplar-birch forest nr. lake shore (crowns and trunks), 50°01’50”N, 95°05’10”E, c. 1150 m a.s.l. GoogleMaps , D. V. Logunov – 11–12/06/1995. PARATYPES: together with the holotype, ZMMU – 2 m 4 f.

Other material examined: RUSSIA: Gorno-Altai Republic: Kosh-Agach Distr., Sailyugem Mt. Range , NW slope, c. 40 km E of Kosh-Agach, c. 20 km NE of Kokorya [=Kokoryu], 50°01’10”N, 89°13’30”E, 2160–2280 m a.s.l., D. V. Logunov, SZMN – 24–25/06/1999, 2 f. Tuva: Kyzyl Distr. , Verkhnii Enisei [=Verkhnii Yenisey, Ulug-Khem] River valley , left riverside, 6–7 km WSW of ‘Bukhara Dzheiran Nursery’ GoogleMaps

Figs 47–52. Male genitalia of Philodromus spp. 47–49. = P. fallax – 47 v, 48 rl, 49 pl. 50–52. = P. angulobulbis – 50 v, 51 rl, 52 pl. Scale bar = 0.1 mm. Specimens: P. fallax : 47–48 = Azerbaijan,

Lenkoran, 49 – E Kazakhstan Region, c. 5 km ESE of Tugyl; P. angulobulbis : the holotype Kyzyl, c. 51°40’25”N, 94°17’30”E, 610–620 m a.s.l., O. V. Lyakhov, SZMN – 19–22/05/1990, 2 m 1 f; Kyzyl Distr., SE foothills of Uyuk Mt. Range, Verkhnii Enisei [=Verkhnii Yenisey, Ulug-Khem] River valley, right riverside, NW vicinities of Kyzyl, 51°43’45”N, 94°24’20”E, 620–700 m a.s.l., D. V. Logunov, SZMN – 17/05/1989, 1 f; Ovyurskii Distr., S foothills of East Tannu-Ola Mt. Range, Irbitei River valley, c. 43 km WNW of Oo-Shynaa, c. 13 km WNW of Ak-Chyraa [=Ak-Chira], 50°44’20”N, 93°08’30”E, 960–1000 m a.s.l., D. V. Logunov, SZMN – 16/06/1995, 1 f; Tandinskii Distr., N foothills of East Tannu-Ola Mt. Range, E shore of Chagytai Lake, 50°01’20”N, 94°46’40”E, 1005–1015 m a.s.l., D. V. Logunov, SZMN – 29/06/1989, 1 f; Tes-Khem Distr., S slope of East Tannu-Ola Mt. Range, c. 4.5 km E of Khol’-Oozhu [=Khol’-Ozhu], Aryskannyg-Khem River canyon, c. 50°45’N, 94°29’E, c. 1200 m a.s.l., Yu. M. Marusik, earlier det. as P. alascensis, SZMN – 16–18/06/1995, 1 f; Erzin Distr., c. 24 km WNW of Erzin, S shore of Bai-Khol’ [=Dus-Khol’] Lake, c. 50°22’25”N, 94°52’05”E, c. 1000 m a.s.l., D. V. Logunov, earlier det. as P. alascensis, SZMN – 31/05/1989, 1 m; Erzin Distr., c. 13 km NW of Erzin, S shore of Dus-Khol’ Lake, c. 50°19’50”N, 95°00’35”E, c. 1040 m a.s.l., D. V. Logunov, SZMN – 06/1995, 2 f; Erzin Distr., SE shore of Tere-Khol’ [=Tore-Khol’] Lake, Eder-Elezin Sands, Sharaa Stand, 50°01’50”N, 95°05’10”E, c. 1150 m a.s.l., S. Koponen & Yu. M. Marusik, ZMTU & SZMN – 11–12/06/1995 1 m 2 f. – KAZAKHSTAN: Pavlodar Area: Pavlodar (town center, Kataeva street), Irtysh River valley, 52°16’50”N, 76°58’20”E, 131–132 m a.s.l., A. V. Gromov, SZMN – 6/06/1998, 1 f. – MONGOLIA: Hövsgöl [=Khubsugul] Aimak, Hövsgöl [=Khubsugul] Lake, Ardagiin-Gol, V. G. Shilenkov, SZMN – 25/07/1977, 1 f.

Diagnosis: Embolus thin. PTA clawlike. VTA small, with the ragged membranous surface. RTA triangular. EA wide, rounded in dorsal view.

Male (the holo- and paratypes): Colouration and pattern as in females ( Fig. 73 View Figs 67–80 ). Measurements (n = 3). Body length 4.70–5.20. PsL 2.15–2.3, PsW 2.15–2.3, OsL 2.55–2.9, OsW 1.65–1.8. Distances between eyes: AME 0.10–0.11, ALE 0.09–0.11, PME 0.08–0.10, PLE 0.08–0.10, AME-AME 0.13, AME-ALE 0.06, PME-PME 0.25, PME-PLE 0.15, MOA-AW 0.31–0.34, MOA-PW 0.41–0.43, MOA-L 0.37–0.42, AME-PME 0.20–0.22, ALE-PLE 0.13–0.14. Length of leg segments: FeI 4.15–4.28, PaI 1.2–1.48, TiI 3.95–4.2, MtI 3.6–4.05, TaI 2.25–2.6, FeII 4.55–4.9, PaII 1.28–1.5, TiII 4.7–4.9, MtII 4.25–4.7, TaII 2.55–2.95, FeIII 3.45–3.65, PaIII 1.15–1.3, TiIII 3.4–3.45, MtIII 2.95–3.25, TaIII 1.75–2, FeIV 3.75–3.85, PaIV 1.05–1.15, TiIV 3.5–3.6, MtIV 3.15–3.45, TaIV 1.7–1.9. Total length of legs: legI 15.15–16.50, legII 17.33–18.95, legIII 12.70–13.65, legIV 13.30–13.85. Spination of leg I: femur: d 0–1–0–1–1; pl 0–0–1–1–1; v -; rl 0–0–1–1–1. tibia: d 0–0–1; pl 1–0–1–0–1; v 2–0–2–0–2ap; rl 1–0–1–0–1. metatarsus: d -; pl 1–0–1–0–1; v; rl 1–0–1–0–1.

Palpus (Figs 27–29). CyL 1.26–1.37, CyW 0.50–0.52, PTiL. 0.31–0.32. Cymbium elongated distally. Embolus arises ventro-medially, with the wide base and the slender apical part, which is slightly bent disto-retrolaterad; embolic tip pointed and curved. The PTA as a large plate with the screwed tip situated retrolaterally of the embolus. The large part of the ST is easily seen in ventral view. The VTA small, with the ragged membranous surface. The RTA large, flat and triangular-shaped (Fig. 28). The proximo-retrolateral part of tibia widened. The palpal patella has a pointed vento-retrolateral projection at its distal end.

Female (the paratypes): Prosoma yellowish brown, with the pattern of reddish and beige stripes ( Fig. 74 View Figs 67–80 ); opisthosoma greyish beige, with the pattern of dark brown and greyish brown stripes. Legs sandy coloured, with dark brown pattern. Measurements (n= 4). Body length 6.3–6.45. PsL 2.15–2.35, PsW 2.15–2.35, OsL 4.05–4.15, OsW 2.9–3.1. Distances between eyes: AME 0.10–0.11, ALE 0.09–0.10, PME 0.08–0.10, PLE 0.08–0.10, AME-AME 0.14–0.15, AME-ALE 0.06–0.07, PME-PME 0.28–0.29, PME-PLE 0.15–0.18, MOA-AW 0.32–0.35, MOA-PW 0.44–0.46, MOA-L 0.37–0.39, AME-PME 0.20–0.22, ALE-PLE 0.17–0.18. Length of leg segments: FeI 3.05–3.4, PaI 1.2–1.25, TiI 2.675–2.95, MtI 2.15–2.55, TaI 1.4–1.65, FeII 3.3–4, PaII 1.05–1.45, TiII 3.2–3.55, MtII 2.65–2.9, TaII 1.65–1.9, FeIII 2.5–3.1, PaIII 1.05–1.15, TiIII 2.25–2.5, MtIII 1.8–2.1, TaIII 1.1–1.25, FeIV 2.8–3.45, PaIV 0.95–1.1, TiIV 2.45–2.75, MtIV 2.05–2.3, TaIV 1.15–1.35. Total length of legs: legI 10.48–11.55, legII 11.85–13.8, legIII 8.7–10, legIV 9.4–10.95. Spination of leg I: femur: d 0–1–0–1–1 or 0–0–1–0–1; pl 0–0–1–1–1; v -; rl 0–0–1–1–1 or 0–0–1–0–1. tibia: d 0–0–1; pl 1–0–1–0–1 or 1–1–0–1–0; v 2–0–2–0–2ap; rl 1–0–1–0–1 or 1–1–0–1–0. metatarsus: d -; pl 1–0–1–0–1; v 2–0–2–0–0ap; rl 1–0–1–0–1.

Epigyne and spermathecae ( Figs 41–42 View Figs 39–46 ). The distal part of CD narrow, the anterior one wide and depressed. The EA singular. The S robust, sac-shaped. The SO is situated anterio-laterally, the ID anteriorly.

Remarks: This species is most similar to P. alascensis . Males can be distinguished by the shape of PTA (clawlike in P. tuvinensis and the flat plate with screwed tip in P. alascensis ; cf. Figs 27, 28 and 24, 25) and by the shape and thickness of embolus (relatively thinner in P. tuvinensis and thicker in P. alascensis ). Females of both species are poorly separable (almost identical), but the shape of the EA is different (cf. Figs 41 and 39 View Figs 39–46 ).

Habitat: Tuva: mountain and semidesert steppes, dry shrub-grass ( Caragana - Poaceae ) steppe, Carex -swamps, and also in crowns and on trunks in river valley poplar-birch forests. Elevations: 130–2300 m a.s.l.

Distribution: N Kazakhstan, the mountain Altai, Tuva and NW Mongolia; for collection localities see Map 3.

A part of the earlier records of P. alascensis (see above) from Tuva ( LOGUNOV et al. 1998, MARUSIK et al. 2000) turned out belong to P. tuvinensis . The same holds true for the records of P. alascensis from China, Xinjiang. Based on the original figures alone ( HU & WU 1989: figs 251.1–4), it is safe to conclude these authors actually dealt with P. tuvinensis .

Etymology: The species is named after the area, from where most of the studied specimens were collected.