Piasites orbitalis, Bordera & Santos, 2022

Bordera, Santiago & Santos, Bernardo F., 2022, A review of Piasites Seyrig (Hymenoptera, Ichneumonidae, Cryptinae), with description of seven new species, Journal of Hymenoptera Research 90, pp. 23-57 : 23

Piasites orbitalis sp. nov.

Figs 2B , 7 , 14D

Diagnosis.

Piasites orbitalis sp. nov. can be distinguished from all other Piasites species by the combination of the following characters: mesoscutum evenly convex, mostly smooth and shiny (Fig. 7C ); mesopleuron dorsally rugose punctate, medially strigose punctate and ventrally coarsely punctate (Fig. 7D ); metapleuron entirely rugose punctate (Fig. 7D ); juxtacoxal carina only present on the anterior part (Fig. 7D ); propodeum strongly rugose; posterior transverse carina weak, forming very low lateral crests, merging in the central part with the strong rugosities of the propodeum (Fig. 2B ); areolet 0.45-0.5 × as high as abscissa of vein 2m-cu above bulla (Fig. 7E ); coxae entirely orange (Fig. 7A, D ); T3 black (Fig. 7A ).

Description.

Female. Fore wing length 7.2-8.0 mm.

Head. In dorsal view, strongly narrowed behind eyes 0.50-0.55 × as long as wide. Gena in dorsal view rounded, weakly swollen. Posterior ocellus separated from eye 1.25-1.40 × its maximum diameter. Distance between posterior ocelli 1.0 × their diameter. Occipital carina complete, weakly v-shaped dorsally. Face slightly convex, mostly smooth and shiny, with sparse punctures and some rugosities in the central part below antennal sockets, middle part delimited by two weak vertical depressions from antennal sockets to clypeal foveae. Clypeal suture present laterally. Clypeus 1.80-1.85 × as broad as medially long, very convex, with relatively long setae in the central part, conspicuously prominent in lateral view, ventral margin slightly rounded with a weak median denticle. Malar space 0.60-0.65 × as long as basal mandibular width. Mandible 1.35 × as long as basal width; dorsal tooth only slightly longer than ventral one. Antenna with 31 flagellomeres, strongly tapered towards apex; flagellomeres from f11 to apex conspicuously flattened ventrally; f1 8.0-8.5 × as long as its maximum width.

Mesosoma. Pronotum mostly strigose punctate, dosally rugose punctate; epomia weak and short, reaching just the anterior depression of pronotum. Mesoscutum 1.20 × as long as wide, evenly convex, predominantly smooth and shiny, with very fine and dense setiferous punctures on median lobe, lateral parts smooth; middle part beetwen posterior end of notaulus strongly rugose; notaulus deep, reaching about 0.7 length of mesoscutum, without transverse keels. Scutellum rugose anteriorly, smooth posteriorly, lateral carinae strong, reaching 0.7 its anterior length. Mesopleuron dorsally rugose punctate, medially strigose punctate and ventrally coarsely punctate; sternaulus deep, with very small transverse keels; epicnemial carina relatively strong, reaching the subtegular ridge. Metapleuron entirely rugose punctate; juxtacoxal carina only scarcely present on the anterior part; submetapleural carina strong and complete. Propodeum strongly rugose; anterior transverse carina complete, centrally narrowly curved towards anterior part; posterior transverse carina weak, forming very low lateral crests, merging in the central part with the strong rugosities of the propodeum. Hind leg with femur about 6.0 × as long as deep. Areolet pentagonal, about 0.7 × as high as wide, relatively small, 0.45-0.5 × as high as abscissa of vein 2m-cu above bulla. Hind wing with vein cu-a 0.4-0.45 × length of first abscissa of CU.

Metasoma. T1 2.1-2.2 × as long as posteriorly broad, granulate, weakly curved dorsally in lateral view, dorso-lateral carina very weak but complete. T2 about 0.8 × as long as posteriorly broad, finely granulate as the following tergites. Ovipositor sheath about 0.55 × as long as hind tibia.

Colour. Body mostly orange. Head black with face and frons except periphery of antennal sockets, ocular orbits (sometimes except an interruption on vertex and malar space), clypeus and labrum, yellow cream; mandible and palpi dark brown; scape, pedicel and f1-3 dark orange, following flagellomeres dark brown, f5-9(10) white. Legs mostly dark orange, all coxae orange. T1 dark orange, anterior half of T2, T3, and anterior part of T4 black; posterior part of T2 and T4 and T5-8 yellow cream. Wings hyaline, slightly yellowish, pterostigma light brown.

Male. Unknown.

Etymology.

The name is a reference to the yellow cream ocular orbits.

Material examined.

2 ♀♀. Holotype: Madagascar • ♀; Rogez, Foret Cote Est; Museum Paris; X-36; A Seyrig; MNHN . Paratype: Madagascar • 1♀; same collection data as for holotype; 1935; MNHN .

Figures

Figure 2. Propodeum of Piasites, dorsal view, females (II): A P. nigricollis sp. nov. (holotype) B P. orbitalis sp. nov. (holotype) C P. lineatus sp. nov. (paratype) D P. seyrigi sp. nov. (paratype). Scale bars: 0.3 mm.

Figure 7. Piasites orbitalis sp. nov., female: A habitus, lateral view (holotype) B head, front view (holotype) C head and mesoscutum, dorso-lateral view (holotype) D head and mesosoma, lateral view (holotype) E front wing, areolet and vein 2 m-cu (paratype). Scale bars: 1 mm (A); 0.5 mm (C, D); 0.3 mm (B, E).

Figure 14. Diagrams showing distribution records for species of Piasites A P. carinatus B P. lineatus sp. nov. C P. nigricollis sp. nov. D P. orbitalis sp. nov.