Platyceroides (Platyceroides) umpquus Paulsen

Paulsen, M. J., 2017, Correction of existing generic and species concepts in Platyceroidini (Coleoptera: Lucanidae: Lucaninae) and the description of four new species of Platyceroides Benesh, Zootaxa 4269 (3), pp. 346-378 : 363-364

Platyceroides (Platyceroides) umpquus Paulsen , new species

Type series. Holotype male ( CNCI) labeled: a) “” So Fork Umpqua R / 14 mi North of / Tiller, Douglas / Co Ore DP Frechin / VI-1 to 5-48 ” ; b) “ Platyceroides / agassizi (LeC.) / Det. by / E. Mengersen 1972”; c) on red paper, “ Platyceroides / umpquus Paulsen / HOLOTYPE ”. Five male, one female paratypes (4 CNCI, 1 MJPC) labeled a, b as holotype. Two male paratypes ( UCDC, MJPC) labeled: a) “ Camas Valley / Ore V. 7.1939”; b) “ A.T. McClay / Collector ”. One male paratype ( CASC) labeled: a) “ Camas Mts. / Coos Co. Or. / VI-17-1911 ” ; b) “ Coll. by / J.R. SLevin”. One female paratype ( CASC) labeled: a) “ Camas Valley / Or . VI-11-36 / F. Lawrence ”; b) “ R. Hopping / Collection ”; c) “[ Platycerus / sp. ♀] / Det. / Ralph Hopping ”; d) “ Platyceroides / agassizi LeC [1978] / Det. B. Gavin”. Two male paratypes ( CASC) labeled: a) “ Myrtle Point / Or. / VI-10-1914 ” ; b) “ Coll. by / E.C. Van Dyke ”; c) “ Van Dyke / Collection ”. Three paratype males (2 CDFA, 1 MJPC) labeled: a) “OR: Douglas County / Tenmile , 5.5 mi. SW / 1 mi. W., VI-14/15 - 99”; b) “ Collector / R.L. Penrose ”. One male paratype ( TAMU) labeled: a) “ Pengra Pass / Ore. VI-9-62 ” ; b) “ Coll. by G.M. / Chamberlain ”; c) “TAMU-ENTO / X0058695”. One male, one female paratype ( MSUC) labeled: “ OREGON: Douglas Co. / Honey Creek / 5 mi E. Idleyld Park / 13 June 1996 G . Parsons ”; b) Platyceroides / agassii (LeConte) / det. G. Parsons 1999”. One male paratype ( USNM) labeled: a) “ OREGON / Brookings / Curry Co. ”; b) “ VI-1963 / NLH Krauss ”. One male paratype ( OSAC) labeled: a) “ OREGON Curry Co. / 4 mi N. Gold Beach / Giesel Mon. Ways. / 14 June 1975 . / Coll. M Gavin”; b) “ Platyceroides / agassizi LeC [1978] / Det. B. Gavin”; c) OSAC barcode label 0 0 0 0 123661. All paratypes with label: on yellow paper, “ Platyceroides / umpquus Paulsen / PARATYPE”.

Description, holotype. Length: 9.8 mm. Width: 4.0 mm. Color: Shiny dark piceous, with indistinct bronze reflection. Head: Form narrow anteriorly, with gena not produced laterally as far as eye. Antennal club moderately large (distinctly shorter than length of scape), size of distal antennomere of club subequal to dorsal surface of eye, antennomeres of club entirely tomentose. Labrum relatively small, shorter than width of distal maxillary palp. Mandibles falcate, externally rounded, internally with 1–2 indistinct teeth. Pronotum: Surface shiny (minutely shagreened) with moderately deep punctures; punctures dense, generally separated by about 1 puncture diameter, distance between punctures becoming greater on center of disc and in explanate lateral margins; median line impunctate. Form broad, lateral margins broadly rounded, posterior angle nearly 90°. Elytra: Surface shiny, with transverse rugae, striate; striae with moderately deep punctures; intervals with punctures finer, irregular. Wings: Wings fully developed. Legs: Mesotibiae and metatibiae not moderately slender as in P. parvicollis and P. aeneus . Abdomen: Male genitalia with permanently everted internal sac sclerotized, elongate, with capitate apex; apex lacking bifurcate process medially, in distal view with 2 strongly sclerotized triangular plates, with opposite side of ostium weakly sclerotized.

Variation in paratypes. Males (n = 17): Length: 9.2–11.5 mm. Width: 4.1–4.6 mm. Color: Light to darker piceous, some with weaker bronze reflection. Pronotum: Surface more or less shiny, shagreened microsculpture obsolete. Posterior angle variable, ± 90°, to rarely produced, subacute. Females (n = 3): Length: 9.6–11.0 mm. Width: 4.2–5.0 mm (body more robust). Color: Lighter piceous to reddish brown, reflection obsolete. Head: Antennal club smaller, about ½ length of scape; not entirely tomentose (mostly glabrous, especially 2 proximal antennomeres). Mandibles internally blade-like. Pronotum: Form larger, longer; posterior margins slightly more than 90°. Elytra: Form more convex. Surface with transverse rugae reduced. Wings: Wings atrophied. Legs: All tibiae distinctly thicker than in males.

Etymology. The specific epithet is derived from the local indigenous word umpqua, meaning ‘across the waters’ or ‘satisfied’, also the name of the Umpqua River and valley of the type locality. The term is Latinized and used as a singular nominative adjective in the masculine gender.

Diagnosis. The apex of the male genitalia ( Fig. 11 B, 11C) is diagnostic for this species because it lacks the bifurcate apex of P. aeneus and the horizontal sclerotized plate visible in distal view in P. californicus . It is darker in color than those species and readily separable by eye from either ( Fig. 11 A).

Distribution ( Fig. 27 ). OREGON: COOS: Camas Mountains; Marshfield; Myrtle Point. CURRY: Brookings; Gold Beach. DOUGLAS: Camas Valley; Honey Creek 5 mi. E Idleyd Park; South Umpqua River 14 mi. N. of Tiller. LANE: Pengra Pass.

Temporal distribution. May (5), June (17). Specimens have been collected from 7 May to 17 June, indicating a short annual period of activity for the adults.

Remarks. The life history of the species is unknown, without any pertinent habitat or host data present on existing labels.

Figures

FIGURES 8 – 11. Platyceroides californicus species group, males: A, dorsal habitus; B, flagellum of male genitalia, lateral view; C, apex of flagellum of male genitalia, ventral view. 8, Platyceroides aeneus, arrow indicates large bifurcate process. 9, Platyceroides californicus. 10, Platyceroides thoracicus, arrow indicates small bifurcate process. 11, Platyceroides umpquus new species.

FIGURES 27 – 28. Distribution of Platyceroides species. 27, Platyceroides californicus species group: P. aeneus (orange triangles); P. umpquus (blue circles); P. californicus (orange squares); P. thoracicus (blue triangles). 28, Platyceroides opacus species group: P. marshalli (blue squares); P. potax (orange squares); P. barrae (blue circles); P. opacus (orange circles).