Platyderus (Eremoderus) insignitus insignitus Bedel, 1902

Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan & Machard, Patrice, 2022, Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini), Deutsche Entomologische Zeitschrift 69 (1), pp. 71-122 : 71

16.1. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902

Figs 4A , 9E , 11D , 15C , 18

Platyderus insignitus Bedel, 1902: 211 (type locality: “Mogador” [= Essaouira], Morocco).

References.

Platyderus insignitus : Bedel 1902: 214; Csiki 1931: 769; Antoine 1957: 235-236; Jeanne 1996: 398; Lorenz 1998: 375; Hovorka and Sciaky 2003: 521; Lorenz 2005: 395; Machard 2017: 102; Machard 2019: 199. Platyderus insignitus insignitus : Hovorka 2017: 756.

Type material.

Five syntypes should be present in MNHN, according to original description. Not examined.

Other material examined.

Morocco: Casablanca-Settat Region : 1♀, ' Marokko (region Doukkala-Abda) / Bir Jdid between El Jadida / and Casablanca / 4.2.2003, leg. M. Egger’ (cWR). Marrakesh-Safi Region: 1♀, 'SW-Morocco, 31.28 N / 09.46W, DIABAT (4 km / S Essaouira), 60 m // J. Kaláb leg., 17.XI.2003 / bushes, sandy dunes, / steppe’ (cASL). Sous-Massa Region: 1♂, 'MAROKKO, s. w. TIZNIT / umg. TIZNIT, 50 m / Strand-AGLOU / I. Puchner, 2.III.2000 ' (cDOS); 1♀, 'SW-Morocco, 29.48N / 09.50W / AGLOU PLAGE, 30 m, / J. Kaláb leg., 26-28.xi.2004 / sandy/grassy places, / half-desert’ (cWR); 1♀, ' MOROCCO, Plage Aglou / NW Tiznit / 29°45'N, 9°54'W / 6 m, 6.V.2011 / lgt. P. Kabátek’ (cWR) .

TME: 5 specimens. TGE: 1♂, 1♀.

Diagnosis.

See “Diagnosis” of P. insignitus presaharensis .

Habitat.

Nothing is known about the bionomics of this species.

Distribution.

According to the available data, P. insignitus insignitus is endemic to Morocco (Fig. 18 ). Antoine (1957: 236) stated that it is a sublittoral species living in the area between Oum er Rbia River to the north and the Anti-Atlas Mountain Range to the south. Machard (2017, 2019) noted that it inhabits the "Littoral atlantique de Casablanca à Agadir". The new data from the environs of Tiznit show that the species occurs south of the city of Agadir as well.

Figures

Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, " Jerusalem Syria ", Jerusalem District, Israel. Scale bars: 1 mm.

Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm.

Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm.

Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13.

Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle).