Perdita (Prockerellia) albonotata Timberlake

Portman, Zachary M. & Griswold, Terry, 2017, Review of PerditasubgenusProcockerellia Timberlake (Hymenoptera, Andrenidae) and the first Perdita gynandromorph, ZooKeys 712, pp. 87-111 : 92

Perdita (Prockerellia) albonotata Timberlake Figures 1A, 2 A–C, 3A, D, 4 A–B, 5A, D, G, 6A, D, G, 7, 8A

Perdita (Procockerellia) albonotata Timberlake, 1954: 403, ♂♀. Holotype female: USA, California, San Bernardino Co., Morongo Valley, 29 September 1944, at flowers of Stephanomeria exigua Nutt. [CAS, type no. 14414]. Examined.

Measurements.

Female (n=10): head width 1.5 mm (1.3-1.8 mm), body length 5.8 mm (5.1-6.8 mm). Male (n=11): head width 1.4 mm (1.2-1.7 mm), body length 5.1 mm (4.0-6.2 mm).

Diagnosis.

Both sexes of P. albonotata have five maxillary palpi and the frons and vertex are weakly tessellate and slightly shining. The female generally has the most extensive facial markings in the subgenus, but the extent is highly variable. In the typical form, the face has the following markings: the clypeus is marked with white on the lateral margins and has a medial white band, the white paraocular marks are broadly triangular and reach the level of the antennal sockets, and the supraclypeal area can be dark or with a pair of white spots (Fig. 5A). The mandible is only slightly expanded on the inner margin and has a long apical tooth (Fig. 5D). The pronotal lobe can be white or dark. The white (sometimes yellowish) abdominal bands on T2-T5 are entire and curve towards the apical margins laterally (Fig. 5G).

The male is unique in Procockerellia due to the simple hind tarsal claws and the presence of an apical nub on the anterior face of the hind tibia, just above the tibial spurs (Fig. 6G). In addition, the male has the pygidial plate broadly and evenly rounded, and the discs of terga have small white markings, with the markings generally more pronounced on the apical terga (Fig. 6D).

Distribution.

Arid regions of the West in Arizona, California, Idaho, Nevada and Utah: Great Basin, southern Colorado Plateau, Mojave Desert (Fig. 8A). There is a single record from the Sonoran Desert. Frequently found in sandy areas.

Phenology.

Floral records.

Asclepiadaceae (1 ♂): Asclepias sp. 1 ♂, Asteraceae (61 ♂ 26 ♀): Asteraceae sp. 1 ♂ 1 ♀, Baileya pleniradiata 2 ♂, Grindelia squarrosa 1 ♂, Helianthus sp. 1 ♀, Isocoma acradenia 1 ♂, Malacothrix sonchoides 3 ♂ 1 ♀, Pectis papposa 2 ♂, Rafinesquia neomexicana 1 ♂, Senecio spartioides 1 ♂, Stephanomeria exigua 3 ♂ 3 ♀, S. pauciflora 2 ♂ 1 ♀, S. sp. 44 ♂ 19 ♀, Brassicaceae (6 ♂ 15 ♀): Streptanthus sp. 6 ♂ 15 ♀, Euphorbiaceae (4 ♂): Croton sp. 1 ♂, C. wigginsii 3 ♂, Fabaceae (1 ♂): Acacia greggii 1 ♂, Loasaceae (1 ♂): Mentzelia multiflora 1 ♂, Malvaceae (1 ♂): Sphaeralcea sp. 1 ♂, Polemoniaceae (1 ♂): Eriastrum wilcoxii 1 ♂, Polygonaceae (3 ♂): Eriogonum cernuum 1 ♂, E. deflexum 1 ♂, E. nummulare 1 ♂.

Additional material examined.

Total specimens: 1403 ♂ 3060 ♀. USA: ARIZONA: Coconino County: Colorado River, Fossil Rapids (36.27333 -112.525): 1 ♂, 28 Sep 2001, T.L. Griswold, J. Sterling, Isocoma acradenia ; Mohave County: Littlefield, N (36.88 -113.92): 1 ♂, 8 May 1998, M. Andres, K. Keen, K. Receveur, C. Schultz; Littlefield (36.88 -113.92): 1 ♂, 15 Jun 1983, W.J. Hanson; Mesquite, 8 mi E (36.81 -113.98): 3 ♂, 26 Apr 1973, F.D. Parker, P.F. Torchio. CALIFORNIA: Riverside County: Palm Springs (33.832 -116.5453): 1 ♂, 2 May 1953, R.M. Bohart; San Bernardino County: (34.868 -115.7731): 2 ♀, 28-29 Apr 2012, collector unknown; 5 ♀, 24-25 May 2012, collector unknown; (34.8794 -115.7808): 3 ♀, 16-17 Aug 2011, collector unknown; 10 ♀, 5-6 Oct 2011, collector unknown; 1 ♀, 23-24 Oct 2011, collector unknown; 1 ♀, 28-29 Apr 2012, collector unknown; 11 ♂ 104 ♀, 24-25 May 2012, collector unknown; Black Canyon (35.1223 -115.394): 2 ♀, 30 May 1998, F.D. Parker; Cedar Canyon (35.16192 -115.44098): 1 ♀, 30 May 1998, F.D. Parker; Hole-in-the-Wall, 2 mi S (35.01783 -115.3843): 7 ♂ 13 ♀, 30 May 1998, F.D. Parker; Kelso, 1 km NE (35.0197 -115.6394): 2 ♂ 3 ♀, 31 May 1998, F.D. Parker. IDAHO: Franklin County: Preston (42.09 -111.87): 1 ♂ 1 ♀, 8 Aug 1972, G.E. Bohart. NEVADA: Churchill County: Sand Mountain, 25 mi SE Fallon (39.3163 -118.4128): 1 ♀, 13 Jul 1980, L. Hanks; Clark County: 0.42 mi E McClanahan Spr. (35.6949 -115.1788): 3 ♂ 1 ♀, 12 May 2004, T.L. Griswold, E. Ahlstrom; 3 ♂ 1 ♀, 25 May 2004, E. Ahlstrom, L. Saul; 1 mi SW Little Virgin Peak (36.5927 -114.21): 3 ♀, 8 Jun 2004, E. Ahlstrom, D. Skandilis; 1.3 mi ESE Mule Spr. (36.0261 -115.5594): 1 ♂, 9 Jun 2004, L. Saul, D. Skandilis; 2.3 mi E Sheep Mtn. (35.7473 -115.2407): 6 ♂ 9 ♀, 12 May 2004, T.L. Griswold, E. Ahlstrom; 3.0 mi SE Rainbow Mtn. (36.0832 -115.4479): 2 ♂, 23 Aug 2004, E. Ahlstrom, Pectis papposa ; 3.5 mi SW Cow Spr. (35.5382 -115.0665): 14 ♂ 1 ♀, 25 May 2004, S.M. Higbee, D. Skandilis; 3.9 mi SSW Whitney Pocket (36.4669 -114.1537): 3 ♂ 7 ♀, 26 May 2005, R. Andrus, S.M. Higbee; Black Mesa, W (36.1607 -114.789): 1 ♂, 9 Oct 1998, T.L. Griswold, Eriogonum deflexum ; Black Ridge, 4.6 mi NW (36.5975 -114.3594): 1 ♂ 1 ♀, 21 Apr 2005, R. Andrus, Stephanomeria pauciflora ; Black Wash (36.4113 -114.0778): 1 ♂, 12 Aug 1998, T.L. Griswold, Baileya pleniradiata ; Bowman Reservoir, E (36.6222 -114.467): 4 ♂ 7 ♀, 5 Aug 1998, M. Andres, C. Schultz; Christmas Tree Pass, W (35.2708 -114.823): 1 ♂, 6 Jun 1998, F.D. Parker; Corn Creek Springs (36.4407 -115.363): 1 ♀, 28 May 1998, F.D. Parker; Eldorado Valley (35.5697 -114.878): 2 ♂, 10 Jun 1998, M. Andres, K. Keen, K. Receveur, C. Schultz; Fire Canyon Wash (36.4548 -114.5088): 18 ♂ 51 ♀, 5 Aug 1998, M. Andres, C. Schultz; Halfway Wash, 1.8 mi NW Virgin River (36.6859 -114.3322): 1 ♂, 6 May 2004, E. Ahlstrom; Highland Range, NW (35.6703 -115.0715): 1 ♂ 1 ♀, 9 Jun 1998, M. Andres, K. Keen; Hot Creek Valley (36.27417 -115.07056): 2 ♂ 2 ♀, 20 Aug 1998, F.D. Parker; Jean Lake, 2.24 mi ENE (35.814 -115.2051): 1 ♂, 10 May 2005, S.M. Higbee, Rafinesquia neomexicana ; Jean Lake, NE (35.8067 -115.2233): 1 ♂ 3 ♀, 8 Oct 1998, T.L. Griswold; Jean, N (35.8102 -115.2998): 1 ♂, 8 Oct 1998, T.L. Griswold; Juanita Springs Ranch, S Riverside (36.6383 -114.2478): 1 ♂, 15 May 1983, F.D. & J.H. Parker; Kyle Canyon (36.3268 -115.3418): 1 ♀, 28 May 1998, F.D. Parker; Las Vegas Dunes (36.286 -114.9667): 1 ♂ 28 ♀, 22 May 1998, M. Andres, K. Keen, K. Receveur, C. Schultz; Las Vegas, NE (36.2798 -115.0355): 1 ♂, 7 Oct 1998, T.L. Griswold; Mesquite (36.8055 -114.0664): 1 ♂, 4 Oct 1988, P.F. Torchio, R.W. Rust, Croton sp.; 14 ♂ 6 ♀, 8 May 1994, P.F. Torchio, D.F. Veirs, S. sp.; Mesquite (36.8144 -114.0703): 3 ♀, 19 Sep 1997, F.D. Parker; Mica Peak (36.3348 -114.1445): 2 ♂ 1 ♀, 7 Jun 1998, T.L. Griswold, F.D. Parker; Mormon Mesa (36.5702 -114.4197): 1 ♂ 1 ♀, 5 Aug 1998, M. Andres, C. Schultz; Mormon Mesa (36.7437 -114.375): 10 ♀, 5 Aug 1998, M. Andres, C. Schultz; Mormon Mesa (36.7447 -114.3778): 36 ♂ 180 ♀, 20 May 1998, M. Andres, K. Keen, K. Receveur, C. Schultz; Mormon Well Rd (36.4355 -115.351): 1 ♀, 15 Sep 1998, W.R. Bowlin; Mormon Well Rd (36.4361 -115.3519): 1 ♂, 15 Sep 1998, W.R. Bowlin; Mormon Well Road (36.534 -115.1067): 6 ♂ 8 ♀, 1 Jul 1998, M. Andres, C. Schultz; Mormon Well Road (36.5492 -115.0995): 6 ♂, 16 Jul 1998, M. Andres, C. Schultz; Mud Wash (36.43 -114.1527): 1 ♀, 19 Sep 1998, W.R. Bowlin; Overton, NE (36.5693 -114.4193): 7 ♂ 5 ♀, 21 May 1998, M. Andres, K. Keen, K. Receveur, C. Schultz; Peek a Boo Canyon (36.5032 -115.1577): 2 ♂ 17 ♀, 16 Jul 1998, M. Andres, C. Schultz; Peek a Boo Canyon (36.5065 -115.1477): 15 ♂ 44 ♀, 16 Jul 1998, M. Andres, C. Schultz; Peek a Boo Canyon (36.515 -115.1327): 8 ♂ 13 ♀, 16 Jul 1998, M. Andres, C. Schultz; Peek a Boo Canyon (36.5325 -115.1128): 12 ♂ 42 ♀, 16 Jul 1998, M. Andres, C. Schultz; Peek-A-Boo Cyn. (36.5522 -115.0991): 1 ♂, 8 Sep 2004, T.L. Griswold, E. cernuum ; Piute Valley (35.46483 -115.052): 8 ♂ 25 ♀, 10 Jun 1998, M. Andres, K. Keen, K. Receveur, C. Schultz;Piute Valley (35.47266 -115.04816): 4 ♂ 1 ♀, 11 Jun 1998, M. Andres, K. Receveur, S. sp.; Pulsipher Wash (36.807 -114.1095): 1 ♂ 2 ♀, 16 Sep 1998, W.R. Bowlin; Pulsipher Wash (36.8073 -114.1143): 4 ♂ 19 ♀, 16 Sep 1998, W.R. Bowlin; Pulsipher Wash (36.8083 -114.1138): 6 ♀, 16 Sep 1998, W.R. Bowlin; Riverside, 4.5 mi SW (36.69137 -114.26056): 1 ♂ 2 ♀, 19 Sep 1997, F.D. Parker; Sandstone Bluffs, E (36.0895 -115.4523): 9 ♂ 24 ♀, 25 Jun 1998, T.L. Griswold; St. Thomas Gap, 0.4 mi E(36.4084 -114.0937): 1 ♂, 20 May 2004, E. Ahlstrom; 109 ♂ 118 ♀, 20 May 2004, S.M. Higbee, E. Ahlstrom; 72 ♂ 100 ♀, 20 May 2004, S.M. Higbee, E. Ahl strom, D. Skandilis, L. Saul; 1 ♂, 28 Jun 2004, E.D. Rentz, Eriastrum wilcoxii ; 2 ♀, 27 Apr 2005, S.M. Higbee; 109 ♂ 387 ♀, 12 May 2005, D. Allen, E. Ahlstrom, R. Andrus, S.M. Higbee; 59 ♂ 193 ♀, 25 May 2005, S.M. Higbee, E. Ahlstrom; 1 ♂, 26 May 2005, S.M. Higbee, E. Ahlstrom; 1 ♂, 8 Jun 2005, D. Allen, S. exigua ; 1 ♂, 8 Jun 2005, S.M. Higbee; 1 ♂, 8 Jun 2005, S.M. Higbee, Acacia greggii ; 2 ♂, 7 Sep 2005, A. Portoluri, C. wigginsii ; 1 ♂, 7 Sep 2005, E. North; 1 ♂, 13 Oct 2005, T.L. Griswold, C. wigginsii ; St. Thomas Gap (36.4023 -114.093): 1 ♂ 4 ♀, 27 May 1998, F.D. Parker; 57 ♂ 59 ♀, 7 Jun 1998, F.D. Parker; 8 ♂ 37 ♀, 12 Aug 1998, M. Andres, T.L. Griswold, C. Schultz; 15 ♂ 46 ♀, 12 Aug 1998, T.L. Griswold, C. Schultz; St. Thomas Gap (36.4041 -114.0933): 1 ♂, 25 May 1998, M. Andres, K. Receveur, C. Schultz, B. pleniradiata ; 68 ♂ 158 ♀, 26 May 1998, M. Andres, K. Keen, K. Receveur, C. Schultz; 1 ♂, 7 Jun 1998, F.D. Parker; 6 ♂ 15 ♀, 8 Jun 1998, T.L. Griswold, Streptanthus sp.; 13 ♂ 54 ♀, 27 Aug 1998, O.J. Messinger, S. Messinger, C. Schultz; 21 ♂ 34 ♀, 6 Oct 1998, T.L. Griswold; 4 ♂ 1 ♀, 6 Oct 1998, T.L. Griswold, S. sp.; St. Thomas Gap (36.4058 -114.0937): 19 ♂, 29 Jun 1998, M. Andres, C. Schultz; 4 ♂, 29 Jun 1998, M. Andres, K. Keen, K. Receveur, C. Schultz; St. Thomas Gap (36.4075 -114.0937): 2 ♀, 4 Aug 1998, M. Andres, C. Schultz; 8 ♂ 5 ♀, 18 Sep 1998, W.R. Bowlin; 6 ♂ 30 ♀, 6 Oct 1998, T.L. Griswold; St. Thomas Gap (36.4083 -114.125): 71 ♂ 63 ♀, 11 May– 12 Jun 1984, R.C. Bechtel, J.B. Knight; Stewarts Point, NW, 9R (36.3853 -114.4123): 1 ♂, 6 May 1998, M. Andres, K. Keen, K. Receveur, C. Schultz; Tramp Ridge, E (36.3905 -114.1287): 2 ♂ 5 ♀, 4 Aug 1998, M. Andres, C. Schultz; Virgin Mountains, W (36.5768 -114.2042): 1 ♂, 6 Oct 1998, T.L. Griswold; Virgin Valley (36.6868 -114.2643): 1 ♂ 10 ♀, 12 Aug 1998, T.L. Griswold, C. Schultz; 7 ♂ 18 ♀, 6 Oct 1998, T.L. Griswold; Whitney Pocket, NW (36.5448 -114.1765): 1 ♀, 26 May 1998, T.L. Griswold; Whitney Pocket (36.5288 -114.1557): 2 ♂ 7 ♀, 27 May 1998, F.D. Parker; Lincoln County: Tule Desert (37.1703 -114.2829): 1 ♂, 17 Aug– 30 Sep 1983, R.C. Bechtel, J.B. Knight; 1 ♀, 15 Sep 1983, R.C. Bechtel, J.B. Knight; Mineral County: Marrietta, 3 mi S (38.2 -118.3): 1 ♀, 16 Aug 1998, F.D. Parker; Nye County: 16 mi E Gabbs (38.8605 -117.623): 26 ♂ 38 ♀, 22 Aug 1998, F.D. Parker; Hot Crk Vly (38.16666 -116.20222): 18 ♂ 19 ♀, 20 Aug 1998, F.D. Parker. UTAH: Cache County: Cornish (41.9756 -111.9525): 2 ♂, 4 Aug 1959, G.E. Bohart, S. sp.; 1 ♀, 4 Aug 1959, G.E. Bohart, R.A. Nielsen, S. sp.; Garfield County: Calf Creek (37.7645 -111.4046): 1 ♂, 18 Jun 2003, S.M. Higbee, S. exigua ; Hole in the Rock Road, Halfway Hollow (37.6338 -111.4449): 7 ♂ 8 ♀, 5-19 Jul 2003, H. Ikerd; 1 ♂, 19-29 Jul 2003, H. Ikerd; Twentyfive Mile Wash (37.5596 -111.3048): 1 ♂ 1 ♀, 18 Sep 2003, A. Johansen, Asteraceae sp.; Kane County: 2.38 mi SE Stave Spr. (37.2344 -112.8769): 5 ♂ 3 ♀, 14-15 Jun 2007, H. Ikerd, K. Davidson; Dry Fork, N (37.441 -111.2307): 1 ♂, 1 Jul 2003, C. Boyers, Mentzelia multiflora ; 1 ♂, 18 Sep 2003, J. Tolliver, S. exigua ; Kitchen Corral Spr., 1.0 mi N (37.2298 -112.1165): 1 ♀, 6 Aug 2003, S.M. Higbee; 1 ♂, 6 Aug 2003, S.M. Higbee, Senecio spartioides ; Paria River W, on HWY 89 (37.12655 -111.95152): 2 ♀, 21 Aug 2008, T.L. Griswold; Sooner Rocks, 0.6 mi WNW (37.333 -111.0713): 3 ♀, 4 Jun 2003, H. Ikerd; Millard County: Oak City (39.37 -112.33): 1 ♀, 24 Jun 1949, G.E. Bohart, Helianthus sp.; Tooele County: 0.5 mi E Wig Mountain (40.3183 -113.0553): 1 ♀, 6 Jul 2005, E. Jarrell, J.S. Wilson; 1 ♀, 6 Jul 2005, J. Wilson; 1 ♂, 15 Aug 2005, J. Wilson, Grindelia squarrosa ; 0.6 mi NW Little Granite Mt. (40.2038 -112.845): 1 ♂, 10 Jun 2003, R. Andrus, S. pauciflora ; 0.7 mi NW Little Granite Mt. (40.2057 -112.8456): 2 ♂ 1 ♀, 10 Jun 2003, O.J. Messinger, S. sp.; 5 ♂ 4 ♀, 10 Jun 2003, O.J. Messinger, R. Andrus; 2 ♂ 2 ♀, 20 Jun 2003, O.J. Messinger, C. Boyers; 3 ♀, 18 Jul 2003, R. Andrus, C. Boyers; 1 ♂, 9 Sep 2003, O.J. Messinger, H. Ikerd; 1 ♂ 4 ♀, 4 Aug 2005, J.S. Wilson, L. Wilson; 17 ♂ 60 ♀, 22 Aug 2005, J.S. Wilson; 17 ♂ 26 ♀, 14 Sep 2005, O.J. Messinger, K.T. Huntzinger; 7 ♀, 29 Sep 2005, K.T. Huntzinger, T.L. Griswold; 1.3 mi W Little Granite Mt. (40.1977 -112.8612): 4 ♂ 3 ♀, 10 Jun 2003, O.J. Messinger, R. Andrus; 1 ♀, 10 Jun 2003, R. Andrus, Malacothrix sonchoides ; 2 ♂ 3 ♀, 20 Jun 2003, O.J. Messinger, C. Boyers; 1 ♀, 1 Jul 2003, J.S. Wilson, O.J. Messinger; 4 ♂, 18 Jul 2003, R. Andrus, C. Boyers; 17 ♂ 33 ♀, 22 Aug 2005, J.S. Wilson; 1 ♂, 14 Sep 2005, K.T. Huntzinger, O.J. Messinger; 10 ♂ 20 ♀, 14 Sep 2005, O.J. Messinger, K.T. Huntzinger; 2 ♂ 5 ♀, 29 Sep 2005, K.T. Huntzinger, T.L. Griswold; 1.8 mi WNW Simpson Butte (40.082 -112.9347): 1 ♂ 6 ♀, 23 Jun 2005, J.S. Wilson, E. Jarrell; 2 mi N Little Granite Mt. (40.225 -112.833): 2 ♂ 3 ♀, 10 Jun 2003, O.J. Messinger, R. Andrus; 1 ♂, 1 Jul 2003, J.S. Wilson, O.J. Messinger; 6 ♂ 24 ♀, 22 Aug 2005, J.S. Wilson; 1 ♂, 14 Sep 2005, O.J. Messinger, K.T. Huntzinger; 2.8 mi NNW Little Granite Mtn. (40.238 -112.8495): 2 ♂, 5 Jul 2005, J.S. Wilson, S. sp.; 1 ♂, 5 Jul 2005, J.S. Wilson, E. Jarrell; 2.8 mi W Simpson Buttes (40.0698 -112.9366): 18 ♂ 5 ♀, 17 Jul 2003, R. Andrus, C. Boyers; 26 ♂ 5 ♀, 30 Jul 2003, J.S. Wilson, C.M. Davidson; 3 ♀, 13 Jun 2005, J.S. Wilson, S. exigua ; 2 ♂ 12 ♀, 13 Jun 2005, J.S. Wilson, E. Jarrell; 4 ♂ 2 ♀, 23 Jun 2005, E. Jarrell, S. sp.; 2 ♂, 23 Jun 2005, J.S. Wilson, M. sonchoides ; 10 ♂ 4 ♀, 23 Jun 2005, J.S. Wilson, S. sp.; 15 ♂ 21 ♀, 23 Jun 2005, J.S. Wilson, E. Jarrell; 1 ♂, 8 Jul 2005, E. Jarrell, S. sp.; 1 ♂, 8 Jul 2005, J. Wilson, E. Jarrell; 6 ♂, 20 Jul 2005, J.S. Wilson, E. Jarrell; 24 ♂ 21 ♀, 22 Aug 2005, J.S. Wilson; 4 ♂ 3 ♀, 30 Aug 2005, O.J. Messinger; 1 ♂ 2 ♀, 15 Sep 2005, K.T. Huntzinger, S. sp.; 1 ♂, 15 Sep 2005, O.J. Messinger, E. nummulare ; 5 ♂ 21 ♀, 15 Sep 2005, O.J. Messinger, K.T. Huntzinger; 1 ♀, 28 Sep 2005, T.L. Griswold, S. sp.; 11 ♂ 33 ♀, 28 Sep 2005, T.L. Griswold, K.T. Huntzinger; 3.5 mi N Wig Mt. (40.3648 -113.088): 5 ♂ 11 ♀, 19 Jun 2003, O.J. Messinger, C. Boyers; 2 ♂ 1 ♀, 17 Jul 2003, R. Andrus, C. Boyers; 4.17 mi SE Wig Mt.(40.2779 -113.0068): 1 ♂, 6 Jul 2005, E. Jarrell; 1 ♂ 1 ♀, 6 Jul 2005, J. Wilson, E. Jarrell; 7 ♂ 3 ♀, 15 Aug 2005, J.S. Wilson, E. Jarrell; 1 ♂, 13 Sep 2005, O.J. Messinger, K.T. Huntzinger; 1 ♂, 26 Sep 2005, T.L. Griswold, K.T. Huntzinger; 4.5 mi SSW White Rock (40.2691 -112.9495): 2 ♀, 1 Aug 2003, J.S. Wilson, C.M. Davidson; 2 ♂, 5 Jul 2005, E. Jarrell, J.S. Wilson; 2 ♂ 1 ♀, 5 Jul 2005, J. Wilson, E. Jarrell; 1 ♂, 5 Jul 2005, J.S. Wilson, Sphaeralcea sp.; 2 ♂ 1 ♀, 20 Jul 2005, J.S. Wilson, E. Jarrell; 4.6 mi WSW Little Granite Mtn. (40.1774 -112.9218): 7 ♂ 12 ♀, 23 Jun 2005, J.S. Wilson, E. Jarrell; 1 ♂, 8 Jul 2005, J. Wilson, E. Jarrell; 1 ♂ 3 ♀, 20 Jul 2005, J.S. Wilson, E. Jarrell; 1 ♂ 14 ♀, 22 Aug 2005, J.S. Wilson; 2 ♀, 30 Aug 2005, O.J. Messinger; 2 ♂ 6 ♀, 15 Sep 2005, O.J. Messinger, K.T. Huntzinger; 2 ♂ 2 ♀, 28 Sep 2005, T.L. Griswold, K.T. Huntzinger; 6.3 mi N Wig Mt. (40.4007 -113.0909): 1 ♂, 21 Jun 2005, J.S. Wilson, E. Jarrell; 1 ♂ 1 ♀, 18 Jul 2005, J.S. Wilson, E. Jarrell; 3 ♂, 15 Aug 2005, J.S. Wilson, E. Jarrell; Camels Back Ridge, 3 mi NNE (40.1705 -112.9314): 1 ♂ 1 ♀, 18 Jul 2003, J.S. Wilson, C.M. Davidson; Camels Back Ridge, 3 mi NNE (40.1706 -112.9297): 1 ♂, 12 Jun 2003, R. Andrus, M. sonchoides ; 13 ♂, 17 Jun 2003, J.S. Wilson; 1 ♂ 5 ♀, 30 Jul 2003, J.S. Wilson, C.M. Davidson; Dugway Proving Grounds, Cedar Ridge, S (site 7) (40.2519 -112.821): 1 ♂, 24 Jun 1997, T. Toler; Dugway Proving Grounds, Dugway, 7 km W (site 6)(40.2333 -112.8313): 4 ♂ 18 ♀, 24 Jun 1997, T. Toler; Dugway Proving Grounds, East Dugway Dunes (40.22111 -112.74361): 1 ♀, 26 Jun 2003, R.L. Johnson; Dugway Proving Grounds, N; Tabbys Peak, 9 km SW (site 13) (40.4296 -113.0946): 10 ♂ 9 ♀, 24 Jun 1997, T. Toler; 3 ♂ 3 ♀, 1 Jul 1997, T. Toler; Dugway Proving Grounds, North Wig Dunes (site 12) (40.22111 -112.74361): 1 ♀, 28 Jul 1997, T. Toler; Dugway Proving Grounds, Wig Mtn., 4.5 km NE (site 8) (40.355 -113.0484): 1 ♂ 2 ♀, 3 Jun 1997, T. Toler; 1 ♂, 24 Jun 1997, T. Toler; Dugway Proving Grounds, Wig Mtn., 8 km E (site 9) (40.2975 -112.9694): 1 ♀, 10 Jun 1997, T. Toler; 1 ♂ 3 ♀, 24 Jun 1997, T. Toler; 2 ♂, 1 Jul 1997, T. Toler; Dugway Proving Grounds, dunes N Wig Mtn. (40.3615 -113.0856): 1 ♀, 18 Jun 2003, R.L. Johnson; 2 ♀, 26 Jun 2003, R.L. Johnson; Dugway Proving Grounds; Dog Area (Ditto), 1 km W (site 19B) (40.1741 -112.9191): 1 ♂ 1 ♀, 3 Jun 1997, T. Toler; Dugway Proving Grounds; Dog Area (Ditto), 8 km N (site 19B) (40.2551 -112.9022): 4 ♂ 10 ♀, 24 Jun 1997, T. Toler; 1 ♂, 1 Jul 1997, T. Toler; Dugway Proving Grounds; Dog Area (Ditto), 8.5 km NE (site 1) (40.2373 -113.8452): 1 ♂ 3 ♀, 24 Jun 1997, T. Toler; Dugway Proving Grounds; Dog Area (Ditto), 9 km N (site 21B) (40.2668 -112.9478): 3 ♂ 3 ♀, 3 Jun 1997, T. Toler; 1 ♀, 24 Jun 1997, T. Toler; Dugway Proving Grounds; Wig Flats, S (site 18) (40.17113 -112.94218): 1 ♀, 3 Jun 1997, T. Toler; Dugway Proving Grounds; Wig Mtn., 10 km WNW (site 12) (40.4 -113.0894): 3 ♂ 6 ♀, 24 Jun 1997, T. Toler; 2 ♂ 1 ♀, 1 Jul 1997, T. Toler; 1 ♂, 24 Jul 1997, T. Toler; Washington County: 1.12 mi WSW Stave Spr. (37.2558 -112.9235): 1 ♀, 7 Jul 2006, B. Hays, F. Nicklen; Firepit Kn., 0.72 mi E (37.3494 -113.0912): 1 ♀, 24 Apr 2007, H. Ikerd, K. Davidson; Firepit Kn., 0.73 mi SE (37.3459 -113.0922): 1 ♀, 26 Jun 2007, K. Davidson; Kolob Terrace Road,.63mi NW, Tabernacle Dome (37.305 -113.1016): 1 ♀, 31 May 2007, H. Ikerd; Oak Creek Cyn. (37.2131 -112.9961): 1 ♂, 8 Jun 2006, F. Nicklen, Asclepias sp.; Paradise Canyon (37.1442 -113.613): 1 ♀, 19 Jun 1983, D. Beck; 1 ♀, 11 Jul 1983, D. Beck; Spendlove Kn., 0.28 mi NNE (37.3404 -113.1059): 1 ♀, 13 Jun 2006, B. Hays, F. Nicklen; 1 ♀, 22 Jun 2006, B. Hays, F. Nicklen; 1 ♀, 13 Jun 2007, H. Ikerd, K. Davidson; St. George, 5 mi N (37.1738 -113.6186): 1 ♀, 22 Jun 1972, F.D. Parker, D. Vincent; Warner Valley; Warner Valley Rd. (37.02541 -113.43376): 25 ♂ 90 ♀, 14-15 May 2012, K. Williams, E. Sadler, D. Denlinger, A. Kelley; 7 ♂ 11 ♀, 15-16 May 2012, K. Williams, E. Sadler, D. Denlinger, A. Kelley; 61 ♂ 372 ♀, 16-17 May 2012, K. Williams, E. Sadler, D. Denlinger, A. Kelley.