Cyrtoscydmini

Jałoszyński, Paweł, 2015, Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 4007 (3), pp. 343-369 : 349-350

Key to Eurasian genera of Cyrtoscydmini ( Protoscydmus Franz omitted)

1. Antenna lacking constriction between antennomeres X and XI which appear closely connected and antennae appear 10-segmented [subtropical Asia and Himalaya Mts]................................... Syndicus Motschulsky ( Figs 34–35 )

- Antennomere X and XI distinctly separated................................................................ 2

2. Antennae nearly as long as body and nearly filiform, very weakly and gradually thickened distally; femur of each leg with narrow longitudinal dorsal groove and small glandular opening [subtropical Asia].................. Loeblites Franz ( Fig. 25 )

- Antennae shorter than body and not filiform, either with club composed of 2–5 terminal antennomeres or distinctly gradually thickened distally; femora lacking dorsal grooves and glandular openings......................................... 3

3. Distance between posterior margin of eye and occipital constriction (i.e., length of temple) comparable to length of eye or larger, eyes not adjacent to anterior margin of pronotum in intact specimens....................................... 4

- Distance between posterior margin of eye and occipital constriction much shorter than length of eye, eyes adjacent or nearly adjacent to anterior margin of pronotum................................................................... 10

4. Each elytron with two basal foveae distinct in dry-mounted and intact specimens.................................. 5

- Each elytron with one basal fovea or rudiments of foveae not visible in dry-mounted and intact specimens.............. 6

5. Mesoventral intercoxal process shorter than mesocoxae and less elevated than mesocoxae (in lateral view its ventral margin concealed by mesocoxae); metaventral intercoxal process with two long spines projecting posteriorly; pronotal base with distinct median pit in addition to lateral pits; head with small pit at posteromesal margin of each supraantennal tubercle; basal ely-

tral foveae connected by U-shaped groove extending anteriorly [subtropical Asia and Himalaya Mts]........................................................................................... Horaeomorphus Schaufuss ( Fig. 23 ) - Mesoventral intercoxal process keel-like, longer than mesocoxae and more elevated than mesocoxae (its ventral margin visible in lateral view, not concealed by mesocoxae); metaventral intercoxal process lacking spines, with straight or concave posterior margin; pronotal base lacking median pit; head lacking pits near supraantennal tubercles; basal elytral foveae not connected [cosmopolitan].............................................................. Euconnus Thomson ( Figs 19–22 )

6. Elytra with basal foveae not visible in dry-mounted specimens [ China: Yunnan]........ Schuelkelia Jałoszyński ( Figs 1–2 )

- Each elytron with one distinct fovea...................................................................... 7

7. Pronotum in dorsal view broadest at or near base and narrowing anteriorly........................................ 8

- Pronotum broadest near middle or in front of middle, narrowing both anteriorly and posteriorly........................ 9

8. Posterior margin of vertex with four distinct tufts or rosettes of dense bristles; maxillary palpomere III weakly narrowing in distal half; maxillary palpomere IV subconical [ Thailand].................................. Siamites Franz ( Fig. 31 )

- Posterior margin of vertex with setae not arranged in paired tufts; maxillary palpomere III strongly narrowing in distal half; maxillary palpomere IV nearly rod-like, very slender and visible as terminal spine [ Malaysia, Indonesia]............................................................................................ Elacatophora Schaufuss ( Fig. 18 )

9. Pronotum elongate, lacking paired basal pits; hypomeral ridges absent; metaventral intercoxal process notched at middle and lacking pair of long spines [Mediterranean].......................................... Leptocharis Reitter ( Fig. 24 )

- Pronotum about as long as broad, with paired basal pits; hypomeral ridges present and complete; metaventral intercoxal process with a pair of robust spines [cosmopolitan]........................ Microscydmus Saulcy & Croissandeau ( Fig. 26 )

10. Head with raised dorsal median triangular area pointing posteriorly and with median longitudinal groove [ China, Nepal]............................................................................. Nanoscydmus Jałoszyński ( Fig. 27 )

- Head lacking raised median triangular area and lacking longitudinal groove..................................... 11

11. Each elytron with one large basal fovea filled with dense setae [panpalaearctic]............ Neuraphes Thomson ( Fig. 28 )

- Each elytron with variously visible but always asetose basal fovea.............................................. 12

12. Pronotum with transverse antebasal groove................................................................ 13

- Pronotum lacking transverse antebasal groove.............................................................. 14

13. Basal elytral fovea large and well-visible; mesoventral intercoxal process posteriorly reaching to middle of mesocoxa; anterior metaventral process present; aedeagus with parameres [panpalaearctic].................. Scydmoraphes Reitter ( Fig. 30 )

- Basal elytral fovea barely noticeable; mesoventral intercoxal process posteriorly reaching posterior margin of mesocoxa; anterior metaventral process absent; aedeagus lacking parameres [Sumatra, Sri Lanka]............................................................................................... Stenichnodes ( Parastenichnaphes Franz ) ( Fig. 32 )

14. Pronotum with rounded lateral margins, lacking lateral edges; metaventral intercoxal process notched but lacking pair of spines; aedeagus with parameres [panpalaearctic].................................... Stenichnus Thomson ( Fig. 33 )

-. Pronotum with lateral edges in posterior half; metaventral intercoxal process with a pair of long spines; aedeagus lacking parameres [ Japan: Shikoku].................................................... Rutaraphes Jałoszyński ( Fig. 29 )

Figures

FIGURES 18 – 35. Representatives of all Eurasian genera of Cyrtoscydmini (except Schuelkelia, illustrated above), dorsal habitus. Elacatophora longipes Jałoszyński (18); Euconnus (s. str.) hirticollis (Illiger) (19); Euconnus (Tetramelus) pubicollis (Müller & Kunze) (20); Euconnus (Napochus) pragensis (Machulka) (21); Euconnus (Pycnophus) impar Sharp (22); Horaeomorphus antennatus Jałoszyński (23); Leptocharis creticus (Pic) (24); Loeblites sabahensis Franz (25); Microscydmus (s. str.) bicavatus Jałoszyński (26); Nanoscydmus baoshanus Jałoszyński (37); Neuraphes (s. str.) elongatulus (Müller & Kunze) (28); Rutaraphes shikokuensis Jałoszyński (29); Scydmoraphes sparshalli Denny (30); Siamites complicatus Jałoszyński (31); Stenichnodes (Parastenichnaphes) ceylonensis (Franz) (32); Stenichnus (s. str.) grebennikovi Jałoszyński (33); Syndicus (s. str.) calcarifer Franz (34); Syndicus (Semisyndicus) leai ranongensis Jałoszyński (35).

FIGURES 1 – 2. Schuelkelia unicornis sp. n., dorsal (1) and lateral (2) habitus of holotype male.