Psallus croceus Van Duzee, 1918: 302

Schuh, Randall T., 2006, Revision, Phylogenetic, Biogeographic, And Host Analyses Of The Endemic Western North American Phymatopsallus Group, With The Description Of 9 New Genera And 15 New Species (Insecta: Hemiptera: Miridae: Phylinae), Bulletin of the American Museum of Natural History 2006 (301), pp. 1-115 : 44-54

Psallus croceus Van Duzee, 1918: 302 (n.sp.).

DIAGNOSIS: Recognized by the weakly pinkish background coloration with conspicuous orange spots covering the entire dorsum (fig. 2), the relatively large size among Ceratopsallus spp. , the average length 3.42, the distinct sexual dimorphism (males with larger eyes than females and cylindrical antennal segment 2), and the structure of the male genitalia, the vesica being moderately long (fig. 15), the phallotheca with a slender spine on the ventral surface, the left paramere with the right-hand prong of the anterior process erect and nearly straight, and the right paramere with the apical processes unequally developed, with the left process being distinctly longer than the right (fig. 15). Most similar to C. pantherinus (Van Duzee) in the large eyes, cylindrical second antennal segment, and the pink-orange spots on the dorsum, but that species with antennal segment 2 much shorter, the vesica much longer, the ventral phallothecal spine flattened, and the right paramere with the apical processes of more or less equal length.

REDESCRIPTION: Male: Moderately large among Ceratopsallus spp. , elongate, nearly parallel-sided, total length 3.15–3.70, length apex clypeus–cuneal fracture 2.19–2.48, width pronotum 0.98–1.19. COLORATION (fig. 1): Body and forewings pinkish; entire dorsum with a moderately dense covering of orange spots; eyes silvery or pale; coloration of appendages pale or weakly orange; membrane weakly to boldly marmorate, with a conspicuous large dark area posterior to the cuneus; veins of membrane pale. SURFACE AND VESTITURE (figs. 2, 13C): As in generic description. STRUC- TURE: Head: Eyes sexually dimorphic, large, vertex relatively narrow (fig. 2), eyes occupying nearly entire height of head; antennal segment 2 cylindrical, of uniform diameter over entire length, very long, length well more that 1.5 times width of head (ratio 1.06:0.73); labium long, surpassing hind trochanters by a short distance. GENITALIA (figs. 15): Vesica: Moderately long, slender, roughly J-shaped. Phallotheca: When viewed laterally bearing a short keel-like ridge on apicodorsal margin, spine on ventral margin slender, relatively long. Parameres: Left paramere with right-hand prong of anterior process very long, erect, nearly straight over much of length, left-hand prong relatively broad curving toward posterior process, posterior process of moderate length, nearly straight in lateral view, curving in dorsal view; right paramere long and broad, apical processes asymmetrically developed with left process much longer than right.

Female: Elongate ovoid; total length 3.03, length apex clypeus–cuneal fracture 2.11– 2.22, width pronotum 0.96–1.01. COLORA- TION (fig. 2): As in male. SURFACE AND VESTITURE: As in male. STRUCTURE: Hemelytra extending just beyond apex of

(AMNH_PBI 00074878).

abdomen; eyes smaller than in male; antennal segment 2 tapered toward base. GENITA- LIA (fig. 15): See generic description.

HOST: Sycamore ( Platanus sp. ) ( Platanaceae ) ( Van Duzee, 1918).

DISTRIBUTION (map 3): Known from Riverside and Alpine Counties, California.

DISCUSSION: Van Duzee (1918) described Psallus croceus from material ‘‘taken on a sycamore tree in Andreas’ Canyon at Palm Springs, Calif.’’ In his description he indicated that the right paramere was ‘‘long, curved and tapered, transverse, reaching across the genital segment’’. He did not mention the tubercle on the left side of the pygophore, although examination of the holotype indicates that it is present.

HOLOTYPE: USA: California: Riverside Co.: Palm Springs [33.83028 ° N 116.54444 ° W], 18 May 1917 – 20 May 1917, E.P. Van Duzee, 1 Oi ( AMNH _ PBI 00077829 ) ( CAS).

PARATYPES: USA: California: Riverside Co.: Palm Springs, 33.83028 ° N 116.54444 ° W, 18 May 1917 – 20 May 1917, E.P. Van Duzee, 3Oi ( AMNH _PBI 00077247– AMNH _PBI 00077249), 2♀ ( AMNH _PBI 00077134, AMNH _PBI 00077135) ( CAS). 1Oi ( AMNH _PBI 00068782) ( USNM).

OTHER SPECIMENS EXAMINED: USA: California: Alpine Co.: Alpine, 38.6 ° N 119.8 ° W, 09 Jul 1929, R.H. Beamer, 2Oi (AMNH_PBI 00074863, AMNH_PBI 00074864) (AMNH). 4Oi (AMNH_ PBI 00074847, AMNH_PBI 00074860–AMNH_ PBI 00074862), 2♀ (AMNH_PBI 00074955, AMNH_PBI 00074956) (KU).

Ceratopsallus pantherinus (Van Duzee) , new combination figures 2, 16; map 3

Psallus pantherinus Van Duzee, 1917: 279 (n.sp.).

DIAGNOSIS: Recognized by the weakly pinkish background with conspicous orange spots covering the entire dorsum (fig. 2), the relatively large size among Ceratopsallus spp. , the average length 3.48, the distinct sexual dimorphism (males with larger eyes than females and cylindrical antennal segment 2), and the structure of the male genitalia, with the vesica being very long compared to other Ceratopsallus spp. , the phallotheca with a flattened spine on the ventral surface, the left paramere with the right-hand prong of the anterior process smoothly curving, and the right paramere with the apical processes more or less equally developed (fig. 16). Most similar to C. croceus (Van Duzee) in the large eyes and the pink-orange spots on the dorsum, but that species with antennal segment 2 much longer, the vesica much shorter, the ventral vesical spine slender, and the right paramere with the apical processes of unequal length.

REDESCRIPTION: Male: Moderately large among Ceratopsallus spp. , elongate, nearly parallel-sided, total length 3.42–3.55, length apex clypeus–cuneal fracture 2.23–2.32, width pronotum 1.05–1.09. COLORATION (fig. 2): Body and forewings weakly pinkish; entire dorsum with moderately dense covering of orange spots; eyes silvery to red in available specimens; coloration of appendages pale or weakly orange; membrane weakly to boldly marmorate, with a large dark area posterior to the cuneus; veins of membrane pale. SURFACE AND VESTI- TURE (figs. 2, 13C): As in generic description. STRUCTURE: Head: Eyes sexually dimorphic, large, vertex relatively narrow (fig. 2), eyes occupying nearly entire height of head; antennal segment 2 cylindrical, of uniform diameter over entire length, relatively short, length about onethird greater than width of head (0.90:0.70); labial apex obscured in all available specimens. GENITALIA (figs. 16): Vesica: Very long, filamentous, forming 2 complete coils (drawn to 50% of scale of congeners). Phallotheca: When viewed laterally with a broadly flattened apex, spine on ventral margin flattened, relatively long. Parameres: Left paramere with relatively short body, right-hand prong of anterior process smoothly curving and tapered toward apex, left-hand prong relatively short and narrow, posterior process relatively short, straight in lateral view, curving in dorsal view; right paramere moderately long and broad, apical processes symmetrically developed, of more or less equal length.

Female: Elongate ovoid; total length 3.02– 3.10, length apex clypeus–cuneal fracture 2.06–2.10, width pronotum 0.92–0.96. COL- ORATION (fig. 2): As in male. SURFACE AND VESTITURE: As in male. STRUC- TURE: Hemelytra just covering abdomen; eyes smaller than in those of male, vertex relatively broad (figs. 2); antennal segment 2 tapered toward base. GENITALIA: Not examined.

HOST: Recorded from Quercus chrysolepis Liebm. and Q. lobata Née (Fagaceae) .

DISTRIBUTION (map 3): Distributed from Josephine County in southern Oregon and south through the northern coastal mountains to the foothills of the southern Sierra Nevada Mountains in California.

DISCUSSION: Van Duzee (1917) indicat- ed that he had described this species on the basis of specimens taken on Quercus agrifolia Née near Cloverdale, Sonoma County , California, and other material taken near Hoberg’s Resort, Lake County, California. He also mentioned having a single specimen from Manitou, Colorado. Van Duzee (1917) did not say whether the Hoberg Resort material was also taken on Quercus agrifolia . I have examined the Manitou, Colorado, specimen; it belongs to Bisulcopsallus polhemorum , n.sp.

HOLOTYPE: USA: California: Sonoma Co. : Cloverdale [38.80546 ° N 123.01722 ° W, 101 m], 03

Aug 1916, E.P. Van Duzee, 1Oi ( AMNH _PBI 00077844) ( CAS).

PARATYPES: USA: California: Lake Co.: Hobergs, near, 38.84351 ° N 122.72443 ° W, 924 m, 02 Aug 1916, E.P. Van Duzee, 1Oi ( AMNH _PBI 00068637) ( CAS). Sonoma Co. : Cloverdale, 38.80546 ° N 123.01722 ° W, 101 m, 03 Aug 1916, E.P. Van Duzee, 1Oi ( AMNH _PBI 00077138), 2♀ ( AMNH _PBI 00077139, AMNH _PBI 00077140) ( CAS).

OTHER SPECIMENS EXAMINED: USA: California: Glenn Co.: 2 mi W of Elk Creek, 39.60027 ° N 122.57407 ° W, 07 Jun 1984, J.D. Pinto, Quercus sp. (Fagaceae) , 4Oi (AMNH_PBI 00083881– AMNH_PBI 00083884) (UCR). Kern Co.: Poso Creek, 35.74596 ° N 119.46998 ° W, 05 Jun 1929, E.P. Van Duzee, 1Oi (AMNH_PBI 00077807), 2♀ (AMNH_PBI 00077811, AMNH_PBI 00077812) (CAS); 05 Jun 1929, R.L. Usinger, 4Oi (AMNH_ PBI 00077806, AMNH_PBI 00077808–AMNH_ PBI 00077810) (CAS). Napa Co.: 2 mi NNE of Angwin, on N side of Howell Mountain, 38.57583 ° N 122.44889 ° W, 396 m, 01 Jun 1978, H.B. Leach, 1Oi (AMNH_PBI 00077250) (CAS). Santa Clara Co. : Los Gatos, 37.22638 ° N 121.97444 ° W, 103 m, 01 Aug 1933, J.A. Kusche, 2Oi (AMNH_PBI 00077251, AMNH_PBI 00077252), 2Oi (AMNH_PBI 00077136, AMNH_ PBI 00077137) (CAS). Trinity Co.: Van Duzen Road, 40.59833 ° N 124.15611 ° W, 05 Aug 1951, Bliven, 1Oi (AMNH_PBI 00077257) (CAS). Tulare Co.: Lemoncove, 36.38278 ° N 119.02361 ° W, 26 Jul 1929, R.H. Beamer, 1Oi (AMNH_PBI 00074953) (KU). Yolo Co.: Davis, 38.545 ° N 121.73944 ° W, 09 Aug 1998, A.G. Wheeler, Jr., Quercus lobata (Fagaceae) , 2Oi (AMNH_PBI 00069412, AMNH_PBI 00069413) (USNM). Oregon: Josephine Co.: 11 miles E Cave Junction, Grayback Campground, 42.16285 ° N 123.43193 ° W, 09 Aug 1979, G. Stonedahl, Quercus chrysolepis (Fagaceae) , 4Oi (AMNH_PBI 00062824–AMNH_ PBI 00062827) (AMNH).

Ceratopsallus pintoi , new species figures 2, 17; map 3

DIAGNOSIS: Recognized by the weakly greenish background with gray brown, relatively small spots covering the hemelytra or the entire dorsum (fig. 2), the moderate size among Ceratopsallus spp. , the average total length 3.00, and the structure of the male genitalia, the vesica moderately long, the phallotheca when viewed laterally with a triangular projection on the apicodorsal margin and a single slender spine ventrally (fig. 17), the left paramere with the left-hand prong of the anterior process long and sinuously curving, the right-hand prong also curving and projecting laterally away from the paramere body, the posterior process of the left paramere of moderate length, nearly straight, the right paramere with the apical processes asymmetrically developed, the left-hand process much longer than right. Similar to C. aquilonius , new species, but apex of phallotheca distinct, C. aquilonius always with a keel-like development rather that a triangular projection on the apicodorsal margin.

DESCRIPTION: Male: Size moderate to large among Ceratopsallus spp. , elongate ovoid, total length 2.50–3.47, length apex clypeus–cuneal fracture 1.76–2.24, width pronotum 0.86–1.02. COLORATION (fig. 2): Body and forewings faded green, often pale in preserved specimens; entire dorsum with moderately dense covering of greenish to brownish spots; eyes silvery to weakly reddish; coloration of appendages pale or greenish; membrane boldly marmorate, with a weakly developed continuous dark area posterior to the cuneus; veins of membrane pale. SURFACE AND VESTITURE (figs. 2, 13C): As in generic description. STRUCTURE: Head: Eyes lacking sexual dimorphism, vertex relatively broad (fig. 2), eyes occupying approximately two-thirds height of head; antennal segment 2 weakly tapered proximally (fig. 2); labium long, reaching hind trochanters, or surpassing them by a short distance. GENITALIA (figs. 17): Vesica: Moderately long, forming one complete loop. Phallotheca: When viewed laterally, with a triangular projection on the apicodorsal margin and a single slender spine on ventral margin. Parameres: Left paramere with left-hand prong of anterior process long and sinuously curving, right-hand prong also curving and projecting laterally away from the paramere body, posterior process of moderate length, nearly straight; right paramere with apical processes asymmetrically developed, the left process much longer than right.

Female: Elongate ovoid; total length 2.52– 3.24, length apex clypeus–cuneal fracture 1.79–2.24, width pronotum 0.85–0.98. COL- ORATION (fig. 2): As in male. SURFACE AND VESTITURE: As in male. STRUC- TURE: Hemelytra extending just beyond apex of abdomen; eyes usually similar in size to those of male (figs. 2); antennal segment 2 tapered toward base. GENITALIA (fig. 17): See generic description.

ETYMOLOGY: Named for John D. Pinto, who during his tenure as a professor of entomology at the University of California, Riverside, collected and documented hosts for this and many other species of Miridae from California and Arizona.

HOST: Recorded from Amelanchier alnifolia (Nutt.) Nutt. , Cercocarpus betuloides Nutt. , C. ledifolius Nutt. , and Purshia tridentata (Pursh.) DC (Rosaceae) , Ceanothus cordulatus Kellogg , C. crassifolius Torr. , C. greggii vestitus (Greene) McMinn , and C. cuneatus (Hook.) Nutt. (Rhamnaceae) , Eriodictyon angustifolium Nutt. (Hydrophyllaceae) , and Quercus sp. (Fagaceae) .

DISTRIBUTION (map 3): Widely distributed from Jackson and Klamath Counties in southern Oregon, south to Los Angeles and Riverside Counties in California, and east to Gila and Mohave Counties, Arizona.

DISCUSSION: Based on available records, this is the most widespread species of

Ceratopsallus , in terms of both geography and host preference. Nonetheless, the male genitalia are remarkably consistent in structure across the entire range of the taxon as here construed.

HOLOTYPE: USA: Arizona: Gila Co. : Old CCC Campground S of Globe on Pioneer Pass Rd [33.39417 ° N 110.78583 ° W], 1433 m, 30 May 1983 – 31 May 1983, R.T. Schuh, G.M. Stonedahl, B.M. Massie, Eriodictyon angustifolium Nutt. (Hydrophyllaceae) , 1Oi ( AMNH _PBI 00063242) ( AMNH).

PARATYPES: USA: Arizona: Gila Co. : Old CCC Campground S of Globe on Pioneer Pass Rd, 33.39417 ° N 110.78583 ° W, 1433 m, 30 May 1983 – 31 May 1983, R.T. Schuh, G.M. Stonedahl, B. M. Massie, Eriodictyon angustifolium Nutt. (Hydrophyllaceae) , 14Oi ( AMNH _PBI 00063240, AMNH _PBI 00063318– AMNH _PBI 00063330), 5♀ ( AMNH _PBI 00063243, AMNH _PBI 00063331– AMNH _PBI 00063334) ( AMNH). Mohave Co.: Hualapai Mountains, SE of Kingman, T20N R15W, 35.18944 ° N 114.05222 ° W, 1585 m, 09 Jun 1983, R.T. Schuh, M.D. Schwartz, G.M. Stonedahl, Quercus sp. (Fagaceae) , 6Oi ( AMNH _PBI 00063302, AMNH _PBI 00063306– AMNH _PBI 00063310), 5♀ ( AMNH _ PBI 00063313– AMNH _PBI 00063317) Ceanothus greggii vestitus (Greene) McMinn (Rhamnaceae) , 8Oi ( AMNH _PBI 00063238, AMNH _PBI 00063239, AMNH _PBI 00063241, AMNH _PBI 00063303– AMNH _PBI 00063305, AMNH _PBI 00063311, AMNH _PBI 00063312), 3♀ ( AMNH _ PBI 00063129, AMNH _PBI 00063244, AMNH _PBI 00063245) ( AMNH). California: Kern Co.: 20 km W of Wofford Heights on Rt 155, 35.70694 ° N 118.6764 ° W, 1500 m, 26 Jul 1999, M.D. Schwartz, Cercocarpus betuloides (Rosaceae) , 2Oi ( AMNH _PBI 00071708, AMNH _PBI 00071709) ( AMNH). Cercocarpus betuloides (Rosaceae) , 1Oi ( AMNH _PBI 00071707) ( CNC). 44.6 km E of jct with Rt 65 on Rt 155, W of Glennville, 35.72889 ° N 118.20954 ° W, 1000 m, 26 Jul 1999, M.D. Schwartz, Cercocarpus betuloides (Rosaceae) , 1Oi ( AMNH _PBI 00071706) ( CNC). 7 km W of Wofford Heights on Rt 155, 35.725 ° N 118.52555 ° W, 1520 m, 26 Jul 1999, Schwartz, Gillespie, Quiring, Ceanothus cordulatus Kell. (Rhamnaceae) , 1Oi ( AMNH _PBI 00071710), 1♀ ( AMNH _PBI 00071711) ( AMNH). Ceanothus cordulatus Kell. (Rhamnaceae) , 1Oi ( AMNH _PBI 00071907) ( CNC). Los Angeles Co.: Glendale, 34.1425 ° N 118.25417 ° W, 12 Jul 1948, E. I. Schlinger, 1Oi ( AMNH _PBI 00074608) ( UCD). Riverside Co.: Menifee Valley, hills on W end, 33.72833 ° N 117.14556 ° W, 549 m, 13 Jun 1979 – 20 Jun 1979, J.D. Pinto, Ceanothus crassifolius (Rhamnaceae) , 22Oi ( AMNH _PBI 00068485, AMNH _PBI 00068486, AMNH _PBI 00082260– AMNH _PBI 00082279), 24♀ ( AMNH _PBI 00082280, AMNH _PBI 00082876– AMNH _PBI 00082880, AMNH _PBI 00082889– AMNH _PBI 00082906) ( UCR); 04 Jun 1979 – 20 Jun 1979, J.D. Pinto, Ceanothus crassifolius (Rhamnaceae) , 5Oi ( AMNH _PBI 00082297– AMNH _PBI 00082301), 4♀ ( AMNH _PBI 00082302– AMNH _PBI 00082305) ( UCR). Shasta Co.: 1 mi W of Fall River Mills, 41.08058 ° N 121.46683 ° W, 1030 m, 07 Jul 1979, R.T. and Joe Schuh, Ceanothus cuneatus (Pursh.) DC (Rhamnaceae) , 15Oi ( AMNH _PBI 00062768– AMNH _PBI 00062781, AMNH _PBI 00063258), 7♀ ( AMNH _PBI 00062782– AMNH _PBI 00062787, AMNH _PBI 00063168) ( AMNH). Brown Butte, 40.83472 ° N 121.52861 ° W, 07 Jul 1947, R.L. Usinger, 4Oi ( AMNH _PBI 00079113– AMNH _PBI 00079116) ( UCB). Cayton, 41.06194 ° N 121.63278 ° W, 19 Jul 1918 – 20 Jul 1918, E.P. Van Duzee, 2Oi ( AMNH _PBI 00077814, AMNH _PBI 00077815), 6♀ ( AMNH _ PBI 00077816– AMNH _PBI 00077821) ( CAS). Tehama Co. : 10 mi W of Mineral, 40.34762 ° N 121.78405 ° W, 27 Jun 1966, O’Brien, 1Oi ( AMNH _PBI 00079112) ( UCB). 12 mi W of Mineral, 40.34756 ° N 121.82208 ° W, 27 Jun 1967, C.W. O’Brien, 4Oi ( AMNH _PBI 00079108– AMNH _PBI 00079111) ( UCB). Tulare Co.: Mineral King Rd E of Three Rivers, 36.47356 ° N 118.8465 ° W, 492 m, 24 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ceanothus cuneatus (Hook.) Nutt. (Rhamnaceae) , det. A. Sanders UCR 140578, 17Oi ( AMNH _PBI 00096933– AMNH _PBI 00096949), 13♀ ( AMNH _ PBI 00096950– AMNH _PBI 00096962) ( AMNH). Oregon: Jackson Co.: 10 mi E of Brownsboro, 42.46833 ° N 122.5129 ° W, 10 Jul 1979, R.T. and Joe Schuh, Purshia tridentata (Pursh.) DC (Rosaceae) , 4Oi ( AMNH _PBI 00062749– AMNH _PBI 00062752), 6♀ ( AMNH _PBI 00062753– AMNH _ PBI 00062757, AMNH _PBI 00063155) ( AMNH). Klamath Co.: 13 mi W of Keno on Rt 66, 42.12667 ° N 122.18228 ° W, 1402 m, 29 Jul 1986, R.T. Schuh, Amelanchier alnifolia (Nutt.) Nutt. (Rosaceae) , 3Oi ( AMNH _PBI 00062758, AMNH _ PBI 00062759, AMNH _PBI 00063257), 2♀ ( AMNH _PBI 00062760, AMNH _PBI 00062761) ( AMNH). 16 mi S of Beatty, 42.21014 ° N 121.26972 ° W, 26 Jul 1957, G.F. Kraft, Cercocarpus ledifolius Nutt. (Rosaceae) , 1Oi ( AMNH _PBI 00076191) ( ORSU). 4 mi NW of Worden on road to Keno, 42.09003 ° N 121.90227 ° W, 17 Jul 1985, G.M. Stonedahl and J.D. McIver, Amelanchier alnifolia (Rosaceae) , 4Oi ( AMNH _PBI 00062762– AMNH _PBI 00062765), 2♀ ( AMNH _PBI 00062766, AMNH _PBI 00062767) ( AMNH).

Ceratopsallus plautus , new species figures 2, 18; map 3

DIAGNOSIS: Recognized by the cream to pale greenish coloration, the brownish spots on the dorsum restricted to the hemelytra and of more or less uniform size and distribution (fig. 2), the relatively small size among Ceratopsallus spp. , the average total length 2.74, and the structure of the male genitalia, the vesica being J-shaped and relatively short, the phallotheca (in common with C. schwartzi and C. vauqueliniae ) bearing two spines, the one on the ventral surface slender, the other on the anterodorsal surface with a broad spoonlike apex, the left paramere distinctly flattened with the righthand prong of the anterior process erect and weakly curving apically, the posterior process of the left paramere being very long, slender, and smoothly curving downward, and the right paramere with the apical processes being more or less equally developed (fig. 18). Most similar to C. schwartzi in the small size, lack of sexual dimorphism, and the form of the male genitalia, but easily distinguished by the slender apicodorsal spine on the phallotheca in C. schwartzi ; distinguished from C. vauqueliniae by the strong sexual dimorphism in the eyes and antennae of that species.

DESCRIPTION: Male: Relatively small among Ceratopsallus spp. , elongate ovoid, total length 2.56–2.87, length apex clypeus– cuneal fracture 1.75–1.90, width pronotum 0.77–0.81. COLORATION (fig. 2): Body and forewings faded green or pale; brownish spots on the dorsum restricted to the hemelytra and of more or less uniform size and distribution; coloration of appendages pale or greenish; membrane boldly marmorate, with a conspicuous and continuous dark area posterior to the cuneus; veins of membrane pale. SURFACE AND VESTI- TURE (figs. 2, 13C): As in generic description. STRUCTURE: Head: Eyes lacking sexual dimorphism, vertex relatively broad (fig. 2), eyes occupying approximately twothirds height of head; antennal segment 2 weakly tapered proximally; labium reaching apex of hindtrochanters. GENITALIA (fig. 18): Vesica: Relatively short, J-shaped. Phallotheca: When viewed laterally with

a weak triangular projection on the apicodorsal margin, a slender spine on the ventral margin, and a second spine on the anterodorsal margin with a broad, spoonlike apex. Parameres: Left paramere distinctly flattened with the right-hand prong of the anterior process erect and weakly curving apically, posterior process of left paramere very long, slender, broadly curving downward; right paramere with apical processes of about equal length.

Female: Elongate ovoid; total length 2.68– 2.92, length apex clypeus–cuneal fracture 1.87–1.98, width pronotum 0.79–0.84. COL- ORATION (fig. 2): As in male. SURFACE AND VESTITURE: As in male. STRUC- TURE: Hemelytra just covering abdomen; eyes usually similar in size to those of male (fig. 2); antennal segment 2 tapered toward base. GENITALIA (fig. 18): See generic description.

ETYMOLOGY: Named for the shape of the anterodorsal spine on the phallotheca. From the Latin, plautus , broad.

HOST: Recorded from Arctostaphylos viscida Parry and Arctostaphylos sp. (Ericaceae) .

DISTRIBUTION (map 3): Known only from Wofford Heights, in the foothills of the Sierra Nevada Mountains, Kern County, California.

HOLOTYPE: USA: California: Kern Co.: 7 km W of Wofford Heights on Rt 155, 35.725 ° N 118.52555 ° W, 1520 m, 26 Jul 1999, Schwartz, Gillespie, Quiring, Arctostaphylos sp. (Ericaceae) , 1Oi ( AMNH _PBI 00071879) ( CNC).

PARATYPES: USA: California: Kern Co.: 7 km W of Wofford Heights on Rt 155, 35.725 ° N 118.52555 ° W, 1520 m, 26 Jul 1999, Schwartz, Gillespie, Quiring, Arctostaphylos sp. (Ericaceae) , 2Oi ( AMNH _PBI 00071796, AMNH _PBI 00071803), 2♀ ( AMNH _PBI 00071814, AMNH _PBI 00071819) ( AMNH). Arctostaphylos sp. (Ericaceae) , 12Oi ( AMNH _PBI 00071793– AMNH _PBI 00071795, AMNH _PBI 00071797, AMNH _PBI 00071798, AMNH _PBI 00071800– AMNH _PBI 00071802, AMNH _PBI 00071804, AMNH _PBI 00071806, AMNH _PBI 00071913, AMNH _PBI 00071914), 14♀ ( AMNH _PBI 00071807– AMNH _PBI 00071813, AMNH _PBI 00071815– AMNH _PBI 00071818, AMNH _PBI 00071880, AMNH _PBI 00071881, AMNH _PBI 00071884) ( CNC). Tulare Co.: Mineral King Rd E of Three Rivers, 36.47356 ° N 118.8465 ° W, 492 m, 24 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Arctostaphylos viscida Parry (Ericaceae) , det. Field ID, 62Oi ( AMNH _PBI 00170512– AMNH _PBI 00170573), 11♀ ( AMNH _PBI 00170574– AMNH _PBI 00170584) ( AMNH).

Ceratopsallus quercicola , new species figures 2, 19; map 4

DIAGNOSIS: Recognized by the cream to weakly orange coloration of the dorsum, the more or less uniform covering of orangebrown spots on the dorsum being restricted to the hemelytra (fig. 2), the small to moderate size among Ceratopsallus spp. , average total length 2.89, and the structure of the male genitalia, the vesica being Jshaped and short (similar in form to C. aquilonius and C. schwartzi ), the phallotheca when viewed laterally with a triangular projection on the apicodorsal margin and a single slender spine ventrally, the left paramere with the branches of the anterior process moderately long, smoothly curving, and forming a crescent, the posterior process relatively short, nearly straight in lateral view, and the right paramere with the apical processes asymmetrically developed, the left process distinctly longer than right (fig. 19). Similar in size and general appearance to C. plautus and C. schwartzi , although coloration in most specimens of C. quercicola with a stronger suggestion of orange (rather than green), and by the structure of the male genitalia, including the left paramere and the phallotheca, with one phallothecal spine in C. quercicola , rather than two as in the other two species.

DESCRIPTION: Male: Size moderate among Ceratopsallus spp. , elongate ovoid, total length 2.83–2.97, length apex clypeus– cuneal fracture 1.88–2.02, width pronotum 0.86–0.93. COLORATION (fig. 2): Body and forewings faded green or pale; brownish spots on the dorsum of more or less uniform size and evenly distributed, either restricted to the hemelytra or covering entire dorsum; coloration of appendages pale or greenish; membrane boldly marmorate, with a weakly developed continuous dark area posterior to the cuneus; veins of membrane pale. SUR- FACE AND VESTITURE (figs. 2, 13C): As in generic description. STRUCTURE: Head: Eyes lacking sexual dimorphism, vertex relatively broad (fig. 2), eyes occupying approximately two-thirds height of head; antennal segment 2 weakly tapered proximally (fig. 2); labium reaching apex of hind trochanters. GENITALIA (fig. 19): Vesica: Relatively short, J-shaped. Phallotheca: When viewed laterally with a triangular projection on apicodorsal margin, apex flattened, a short, slender spine on ventral margin. Parameres: Left paramere with branches of anterior process smoothly curving, more or less lying in a single plane, posterior process relatively short, nearly straight in lateral view; right paramere with the apical processes unequally developed, the left process being distinctly longer than the right.

Female: Elongate ovoid; total length 2.64– 2.87, length apex clypeus–cuneal fracture 1.79–1.98, width pronotum 0.87–1.08. COL- ORATION (fig. 2): As in male. SURFACE AND VESTITURE: As in male. STRUC- TURE: Hemelytra extending just beyond apex of abdomen; eyes usually similar in size to those of male (fig. 2); antennal segment 2 tapered toward base. GENITALIA: Not examined.

ETYMOLOGY: Named for its habit of feeding on at least one species in the genus Quercus Linnaeus (Fagaceae) , among other plant groups.

HOSTS: Recorded from Arctostaphylos pungens Kunth , Arctostaphylos sp. (Ericaceae) , and Quercus oblongifolia Torr. (Fagaceae) .

DISTRIBUTION (map 4): Known from moderate elevations in Riverside County, California, and from Cochise and Pima Counties in southern Arizona.

HOLOTYPE: USA: Arizona: Pima Co. : 7.5 mi S of Coronado Natl. Forest boundary on Mount Lemmon Rd [32.31 ° N 110.72 ° W], 1433 m, 11 Jun 1983, Schuh, Schwartz, and Stonedahl, mercury vapor light, 1Oi ( AMNH _ PBI 00063251 ) ( AMNH).

PARATYPES: USA: Arizona: Cochise Co. : 2.9 mi on Rt 92, Carr Canyon, Huachuca Mountains, 31.45382 ° N 110.25813 ° W, 3 Jun 1997, T.J. Henry and A.G Wheeler, Jr., Arctostaphylos pungens (Ericaceae) , 5Oi ( AMNH _PBI 00068462– AMNH _PBI 00068466) ( USNM). Ash Canyon Road, 0.5 mi W of Hwy 92, 31.38194 ° N 110.22444 ° W, 1554 m, 31 May 1993, N. McFarland, 3Oi ( AMNH _PBI 00068467– AMNH _PBI

00068469) ( USNM); 4 Jun 1997, T.J. Henry and A.G Wheeler, Jr., Arctostaphylos sp. (Ericaceae) , 2Oi ( AMNH _PBI 00068470, AMNH _PBI 00068471) ( USNM); 11 Jun 1997, T.J. Henry and A.G. Wheeler, Jr., 1Oi ( AMNH _PBI 00068474) ( USNM); 2 Jun 1997, T.J. Henry and A.G. Wheeler, Jr, 2Oi ( AMNH _PBI 00068476, AMNH _PBI 00068477) ( USNM). Huachuca Mountains, Carr Canyon, 31.45667 ° N 110.23889 ° W, 3 Jun 1997, T.J. Henry and A.G Wheeler, Jr., Arctostaphylos pungens (Ericaceae) , 7Oi ( AMNH _PBI 00063250, AMNH _PBI 00068459– AMNH _PBI 00068461, AMNH _PBI 00068472, AMNH _PBI 00068473, AMNH _PBI 00068475) ( USNM). Pima Co. : 7.5 mi S of Coronado Natl. Forest boundary on Mount Lemmon Rd, 32.31 ° N 110.72 ° W, 1433 m, 11 Jun 1983, Schuh, Schwartz, and Stonedahl, Quercus oblongifolia (Fagaceae) , 39Oi ( AMNH _ PBI 00063246– AMNH _PBI 00063249, AMNH _ PBI 00063335– AMNH _PBI 00063347, AMNH _ PBI 00063354– AMNH _PBI 00063375), 34♀ ( AMNH _PBI 00062739– AMNH _PBI 00062748, AMNH _PBI 00063252– AMNH _PBI 00063256, AMNH _PBI 00063348– AMNH _PBI 00063353, AMNH _PBI 00063376– AMNH _PBI 00063388) ( AMNH). California: Riverside Co.: San Jacinto Mountains, Pinon Flat, 33.81444 ° N 116.67833 ° W, 28 May 1940, C.D. Michener, Arctostaphylos sp. (Ericaceae) , 2Oi ( AMNH _PBI 00079772, AMNH _PBI 00079773) ( UCB).

Ceratopsallus ribesi (Knight) , new combination figures 2, 20; map 4