Pseudeurybata browni, Marshall, S. A., 2016

Marshall, S. A., 2016, Revision of the genus Pseudeurybata Hennig (Diptera, Micropezidae, Taeniapterinae), Zootaxa 4132 (2), pp. 254-268 : 258-260

Pseudeurybata browni new species

Figs. 8–16

Description. 16–18mm. Head with epicephalon shining but otherwise pollinose. Frontal vitta broad, barely distinguishable from narrow orbital strips anterior to ocelli, part posterior to ocelli about half as wide, margin slightly concave, posteriormost part slightly expanded. Two lower fronto-orbital setae. Base of arista yellow and slightly swollen; arista bare. Clypeus bare and shining.

Thorax mostly orange, sides of mesonotum above postpronotal lobe darkened due to a higher density of microsetulae; postpronotal lobe relatively pale. Anepisternum and anterior part of katepisternum with very sparse small dark setae.

Legs black to dark brown except for orange coxae and femoral bases, hind femur orange on basal third. Fore tarsomere one with an inconspicuous ventral fringe of brown setulae. Wing infuscated, halter knob orange.

Abdomen of female with pleuron cream yellow to white ventrally, brown on dorsal half or more. Female oviscape broad and brown basally, abruptly tapered and black distally, 4X length of T6. Spermathecal ducts smooth until spermathecal stem, paired spermathecae very elongate, sausage-shaped, sparsely but prominently tuberculate; single spermatheca similar but much smaller.

Male abdomen with pleuron 2 swollen and pigmented similarly to female abdomen, pleura 2–5 largely black (but available material shrivelled with pigmentation hard to interpret). Genital fork of sternite 5 simple, with inner basal processes forming small, spinulose, almost sessile lobes and with arms almost straight (slightly converging) and parallel-sided bearing 6–7 small inner teeth. Distiphallus broad, terminating in a dense phallic bulb half as long as basal part of phallus.

Type material (all Costa Rica). Holotype (♂, MNCR). Puntarenas, San Luis Monteverde, 1000–1350m, Feb.1994, Z. Fuentes. Paratypes. Alajuela, San Ramon Biol. Res, 900m, Feb–Mar, May.2000, P. Hanson (5♀, DEBU); Upala, Sector San Ramon de Rios, 27.Feb–27.Abr., 620m, Malaise (♀, MNCR); Guanacaste, near Turin, 24.Feb.1960, at light, Charles Palmer (♂, SEMC); ibid, 7.April (♀, SEMC); Puntarenas, same as holotype but April and January.1994, (♀,♂, MNCR); Monteverde "con Pseudoscorpion" 13.v.2000 (1♀, MNCR); Monteverde, 14.June.2000, M. Buck (♂, DEBU); Buen Amigo, San Luis Monteverde, A.C. Arenal, 1000–1350m, Malaise trap, April.1994, Z. Fuentes (6♀, MNCR); Santa Elena, San Luis Lodge, on dung, Feb.2002, P.D. Careless (1♀, DEBU).

Etymology. The specific name of Pseudeurybata browni is a patronym honoring dipterist Brian Brown, and incidentally reflective of the diagnostically uniform brown to black legs of this species. Distribution. Costa Rica.

Remarks. Pseudeurybata browni is unusual amongst congeners in having the legs uniformly dark distal to the base of the femur, and it is the only species other than P. stigmata to have an orange halter knob.

Figures

FIGURES 8 – 16. Pseudeurybata browni. 8, habitus, paratype female, lateral view; 9, front of head to show clypeus, lateral view; 10, head and anterior thorax, lateral view; 11, head and anterior thorax, dorsal view; 12, spermathecae and associated structures, dorsal view; 13, male genital fork, ventral view; 14, male terminalia, lateral view; 15 apex of male abdomen, lateral view; 16, apex of distiphallus to show phallic bulb, lateral view.