Pseudolycoriella raki, Köhler, 2019

Köhler, Arne, 2019, The genus Pseudolycoriella Menzel & Mohrig, 1998 (Diptera, Sciaridae) in New Zealand, Zootaxa 4707 (1), pp. 1-69 : 34-37

Pseudolycoriella raki sp. n.

( Fig. 41 )

urn:lsid:zoobank.org:act:NomenclaturalActs/ 01500E3C-735A-400F-968D-3B3E1F4A14C9

Literature. Pseudolycoriella jejuna (Edwards, 1927) : Mohrig & Jaschhof (1999): 37 [misidentification].— Köhler & Mohrig (2016): 108 [misidentification].

Material studied. Holotype male. New Zealand: North Island, Far North , Waipoua Forest , alongside Highway 12, altitude 300 m, mixed mature Kauri /podocarp forest, Malaise trap, 24.07– 17.08.2001, leg. M. & C. Jaschhof ( NZAC, SDEI-Dipt-0000834) . Paratypes. 3♂♂ same locality and same date as holotype (1x NZAC; 2x SDEI) . 1♂ North Island, Far North , Puketi Forest , Waipapa River Valley , mixed secondary wood of Kauri and Podocarpus with tree fern, sweep net, 01– 02.01.1993, leg. M Jaschhof ( PWMP [previously misidentified, published as Psl. jejuna in Mohrig & Jaschhof 1999]) . 1♂ North Island, Thames-Coromandel , Coromandel Range , 5 km east of Coromandel , dense Podocarpus secondary wood with tree ferns, sweep net, 28.12.1992, leg. M. Jaschhof ( PWMP [previously misidentified, published as Psl. jejuna in Mohrig & Jaschhof (1999)]) . 1♂ North Island, Whakatane-Distrikt , White Pine Bush 3 km southwest of Whakatane , Podocarpus wood, sweep net, 26.12.1992, leg. M. Jaschhof ( PWMP [previously misidentified, published as Psl. jejuna in Köhler & Mohrig 2016]).

Description. Male. Head. Head capsule brown. Eye bridge three to four facets wide. Scape and pedicel bright yellow, strongly contrasting with the brown flagellum; fourth flagellomere 2.6–3.4 times longer than wide; necks of flagellomeres well differentiated, paler than body of flagellomeres, flagellomere necks at antenna apex show distal dark rings; surface of flagellomeres rough with deep pits, sensilla of two different lengths, small ones and longer, curved sensilla; setae on the flagellomere longer than the flagellomere width, slightly curved, arising from small elevations; apical flagellomeres very rough with prominent elevations. Maxillary palp very long and three-segmented, first or third palpomere longest, second one shortest; first palpomere with long sensilla and four to five long bristles, one bristle longer and more robust, located on the outer side. Prefrons and clypeus bulging. Thorax pale brown, slightly paler than head, with some indistinct lateral brightening. Posterior pronotum bare. Anterior pronotum with three to eight setae. Episternum 1 with two to eight setae. Mesonotum with four to five robust lateral bristles. Scutellum with three to four robust bristles and several minor setae. Katepisternum as long as high. Wing. Length 2.2–2.9 mm; width/length ratio 0.38–0.41. Membrane transparent and without macrotrichia, anal area present; all posterior veins distinct, except faint stem of M, apical fifth to two-fifths of R 5 with macrotrichia on ventral and dorsal side; bM and r-m bare; R 1 0.8–1.0 times as long as R; c/w ratio 0.74–0.79; r-m 0.8–1.2 as long as bM. Haltere brownish; head of haltere slightly longer than shaft. Legs yellowish pale brown, paler than thorax. Tibial organ surrounded by a circular border and wider than half of the width of the tibial apex; tibial organ consists of an irregular row of bristles. Fore tibia with one robust bristle among the vestiture, or without. Mid tibia with one to three robust bristles among the vestiture. Posteriodorsal row of seven to eleven bristles occupies half to two-thirds of the apex of the tibia. All tibial spurs equal in length. Claws with robust teeth. Abdomen concolourous with thorax, with long, dark setae. Hypopygium ( Fig. 41 ). Gonocoxites wider than long, inner side of gonocoxites with small setae, outer side with long setae; ventral, inner side of gonocoxites basally separated. Gonostylus curved and slender, 2.4–2.6 times longer than wide, apex not clearly separated, apical end broad, blunt, on ventral side with a huge apical cavity scattered with few microtrichia, one spine and one whip-lash hair present, inner base of gonostylus without or only with a minor angle. Tegmen of the typical ground plan of the Psl. jejuna complex, onion-shaped but more elongated than in Psl. jejuna ; dorsal folds present; parameral apodeme basally strongly sclerotized, distal branches forming a triangle. Area of teeth absent. Ejaculatory apodeme dark, broad and long; base of ejaculatory apodeme delicate, long, u-shaped. Posterior portion of gonocoxal apodeme medium sized and brown, joined medially. Anterior portion of gonocoxal apodeme long and dark brown, widely separated.

Body size: 2.3–2.9 mm.

Female. Unknown.

Genetic distances. Only one specimen was successfully sequenced. For the COI sequence, the nearest neighbour is Psl. tuakana , diverging by a minimum of 8.42%. For the 28S sequence the nearest neighbour is also Psl. tuakana , diverging by a minimum of 0.44%.

Etymology. Raki is the Māori word for northern.

Distribution. New Zealand.

Discussion. See discussion paragraph for Psl. dagae .

Figures

FIGURES 37–42. Pseudolycoriella jejuna complex, hypopygia. 37. Pseudolycoriella sudhausi sp. n. 38. Pseudolycoriella whena sp. n. 39. Pseudolycoriella porotaka sp. n. 40. Pseudolycoriella nahenahe sp. n. 41. Pseudolycoriella raki sp. n. 42. Pseudolycoriella dagae sp. n. Abbreviation: ba = basal angle.