Psyra angulifera (Walker, 1866)

Liu, Zulian, Xue, Dayong, Wang, Wenkai & Han, Hongxiang, 2013, A review of Psyra Walker, 1860 (Lepidoptera, Geometridae, Ennominae) from China, with description of one new species, Zootaxa 3682 (3), pp. 459-474 : 461-464

Psyra angulifera (Walker, 1866)

Figs 1–2 , 25 , 40 , 53 , 64

Scotosia angulifera Walker, 1866 , List Specimens lepid. Insects Colln Brit. Mus., 35: 1687. Holotype 3, India: North Hindostan (BMNH).

Psyra angulifera: Prout, 1920 , in Seitz, Macrolepid. World, 4: 410, pl. 24: g.

Material examined. INDIA ( BMNH): 13 (Holotype), North India, BMNH (E)#1008371 ( Geometridae genitalia slide no. 11632); 13, Darjiling, F. Möller, Rothschild Bequest B.M. 1939-1, BMNH (E)#1008372. CHINA: Hubei ( IZCAS): 1Ƥ, Shennongjia, 950 m, 18.VII.1980. Sichuan ( IZCAS): 13, Emei Shan, Jiulaodong, 1800–1900 m, 28.VII.1952, coll. Huang Keren; 13, same locality, 6.VII.1952, coll. Wang Zongyuan; 1Ƥ, Emei Shan, Qingyinge, 800–1000 m, 21.VI.1959, coll. Huang Keren; 13, Pingwu, Laohegou, Caiyuchang, 1630 m, 14.VIII.1999, coll. Zhou Xin (slide no. Geom-1997); 43, Yingjing, Siping, 1100 m, 24.VI.1984, coll. Liu Dajun (slide no. Geom- 1989). Yunnan ( IZCAS): 13, Tengchong, Dahaoping, 2020 m, 24–26.V.1992, coll. Xue Dayong; 13, Tengchong, Heinitang, 1930 m, 28–30.V.1992; 13, Cangyuan, Fabao, 790 m, 21.V.1980, coll. Gao Ping; 13, Cangyuan, 1100 m, 16.V.1980, coll. Gao Ping. Tibet ( IZCAS): 1Ƥ, Zham, 20.VII.1984, coll. Hu Shengchang (slide no. Geom- 1991).

Diagnosis. P. angulifera is distinguished by the clear golden brown borders to the black patches on the forewing, though its wing pattern is similar to P. szetschwana . These two species can be distinguished by the following characters: the ground colour of P. angulifera is much darker than that of P. szetschwana ; the medial line on the hindwing in P. angulifera is not as distinct as in P. szetschwana . In the male genitalia, P. angulifera shares the characters of very narrow costal basal lobe and valva with P. szetschwana ; the valva is almost straight in P.

angulifera but the tip is curved in P. szetschwana ; the tip of the costal basal lobe is tapering in P. angulifera but swollen in the latter; in addition, the saccus is not expanded in P. angulifera but projects as a triangle in P. szetschwana ; the aedeagus has two bundles of spines as in P. cuneata , P. szetschwana , P. gracilis , P. spurcataria and P. crypta , but bears more spines; the cornutus is less pointed and slender than in most other species, but as broad as in P. bluethgeni and P. breviprotrusa . In the female genitalia, the antrum and ductus bursae are broader than in most other species; the corpus bursae is much bigger and longer than the ductus bursae and antrum, while in other species, except for P. bluethgeni , the corpus bursae is quite small.

Distribution. China (Hubei, Sichuan, Yunnan, Tibet), India, Nepal.

Figures

FIGURES 1 – 24. Adults of Psyra. 1 – 2. P. angulifera. 1, male (holotype, BMNH); 2, male, from Sichuan (IZCAS); 3 – 5. P. boarmiata. 3, male (holotype? of P. boarmiata boarmiata, MNHU); 4, ditto, female, from Jilin (IZCAS); 5, male (holotype of P. boarmiata subcuneata, BMNH); 6 – 8. P. cuneata. 6, male (holotype of P. cuneata cuneata, BMNH); 7, male, from Yunnan (IZCAS); 8, female (P. cuneata matsumurai, BMNH); 9 – 11. P. szetschwana stat. nov. 9, male (syntype of P. cuneata szetschwana, ZFMK); 10, male (syntype of P. cuneata lidjangica, ZFMK); 11, male, from Yunnan (IZCAS); 12 – 13. P. dsagara stat. nov. 12, male (lectotype, ZFMK); 13, male, from Hunan (IZCAS); 14, P. conferta, male (holotype, BMNH); 15, P. moderata, male (holotype, BMNH); 16, P. gracilis, male, from Yunnan (IZCAS); 17, P. similaria, male (syntype, BMNH); 18 – 19. P. spurcataria. 18, male (lectotype, BMNH); 19, male, from Yunnan (IZCAS); 20, P. rufolinearia, female (holotype, BMNH); 21 – 22. P. falcipennis. 21, male, from Hubei (IZCAS); 22, female, from Fujian (IZCAS); 23, P. breviprotrusa sp. nov., male (holotype, IZCAS); 24, P. bluethgeni, male (syntype, MNHU). Scale bar = 1 cm.

FIGURES 25 – 32. Male genitalia of Psyra. 25, P. angulifera; 26, P. boarmiata boarmiata; 27, P. boarmiata subcuneata (holotype, BMNH slide no. 15920); 28, P. cuneata cuneata; 29, P. szetschwana (syntype, ZFMK, Wehrli slide no. 5306); 30, ditto; 31, P. dsagara; 32, P. conferta (paratype, BMNH, Inoue slide no. 6040). Scale bar = 1 mm.

FIGURES 40 – 52. Aedeagus of Psyra. 40, P. angulifera; 41, P. boarmiata boarmiata; 42, P. cuneata cuneata; 43, P. szetschwana; 44, P. dsagara; 45, P. c o n f e r t a (paratype, BMNH, Inoue slide no. 6040); 46, P. m o de ra t a (holotype, BMNH slide no. 16391); 47, P. gracilis; 48, P. similaria (from Dr. Yazaki); 49, P. spurcataria; 50, P. falcipennis; 51, P. breviprotrusa (holotype); 52, P. bluethgeni (BMNH, Inoue slide no. 8325). Scale bar = 1 mm.

FIGURES 53 – 63. Female genitalia of Psyra. 53, P. angulifera; 54, P. boarmiata subcuneata; 55, P. cuneata cuneata; 56, P. cuneata matsumurai (BMNH, Inoue slide no. 9507); 57, P. szetschwana; 58, P. dsagara; 59, P. conferta (paratype, BMNH, Inoue slide no. 9509); 60, P. m o d e r a t a; 61, P. si m il a r i a; 62, P. spurcataria; 63, P. falcipennis. Scale bar = 1 mm. 64 – 72. Signa of female genitalia of Psyra. 64, P. angulifera; 65, P. boarmiata subcuneata; 66, P. cuneata cuneata; 67, P. szetschwana; 68, P. dsagara; 69, P. m o d e r a t a; 70, P. similaria; 71, P. spurcataria; 72, P. falcipennis. 73. Sternite 8 of male abodomen of P. spurcataria. Scale bar = 0. 1 mm.