Ptomaphaminus kinabatanganensis Njunjic , Schilthuizen & Perreau

Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva, 2018, A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae), ZooKeys 777, pp. 57-108 : 84

Ptomaphaminus kinabatanganensis Njunjic, Schilthuizen & Perreau sp. n. Figures 11d, 12u, v, 13m, n

Material.

Holotype: Malaysia, Sabah, Lower Kinabatangan, Batu Batangan, Gua Kolam, 5°27.557'N, 118°06.118'E, 24. ii– 1.iii.2016 (leg. I. Njunjić et al., BOR/COL/14218), 1 male. Paratypes: Sabah. Lower Kinabatangan, Batu Batangan, Gua Babi, 5°27.570'N, 118°06.088'E, 24-28.ii.2016 (leg. field course students, BOR/COL/14219-14220), 2 individuals; Lower Kinabatangan, Gua Fico, 5°27.135'N, 118°08.769'E, 18.iii.2015 (leg. I. Njunjić et al., CMPR), 1 female; Lower Kinabatangan, Batu Batangan, Gua Babi, N 5°27.570'N, 118°06.088'E, 24-28.ii.2016 (leg. I. Njunjić et al., CMPR), 1 female; Lower Kinabatangan, Batu Batangan, Gua Ikan, 5°27.558'N, 118°05.891'E, 18.iii.2015 (leg. I. Njunjić et al., CMPR), 2 males.

Description.

Length: 1.45-1.60 mm. General colour brown; antenna, mouthparts, and protarsi yellowish, the other tarsi light brown. Winged. Eyes well developed. Pronotum 1.55 times wider than long. Elytra 1.25 times longer than wide (slightly wider in males than in females). Female sutural apex of elytra angular but not protruding backwards. Male protarsi 0.65 times as wide as the apex of protibia. Spiculum gastrale of the male genital segment shortly protruding beyond the apex of epipleurites, significantly dilated (Figure 13m). Aedeagus approximately 4 times smaller than the body length, parallel on the first third of its length, then regularly narrowed, the sides linearly convergent towards the apex in dorsal view (Figure 12u), straight in lateral view (Figure 12v). Apex of the median lobe with a long hook deflexed towards the ventral side, but not retroverted in lateral view (Figure 12v). Six lateroventral preapical setae on each side. Internal stylus of the endophallus long and moderately sinuate. Female genital segment with reduced gonocoxites. Spermiduct not helical. Spermatheca rounded at the apex without sclerotised plate (Figure 13n).

Differential diagnosis.

Female with spiniform elytra; male with long apical hook of the (relatively short but straight) aedeagus, which, however, is not retroverted.

Habitat and distribution.

Known exclusively from three caves in the lower Kinabatangan valley: Gua Babi, Gua Fico, and Gua Ikan.

Etymology.

Named after the lower Kinabatangan valley, in which the specimens were collected.