Quedius (Raphirus) illyricus, Wendeler, 1928

Assing, Volker, 2018, On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae), Linzer biologische Beiträge 50 (1), pp. 149-182 : 159-160

Quedius (Raphirus) illyricus W ENDELER, 1928 ( Map 2 )

Quedius albanicus BERNHAUER, 1926: 267 f.; preoccupied.

Quedius illyricus WENDELER, 1928: 298 ; replacement name.

Quedius paganettii BERNHAUER, 1936:: 308 f.; nov.syn.

Quedius schipkanus SCHEERPELTZ, 1937: 223 ff.; nov.syn.

Quedius pseudopyrenaeus COIFFAIT, 1967: 396 f.; nov.syn.

T y p e m a t e r i a l e x a m i n e d: Q. albanicus: Lectotype ♂, present designation: " ♂ / Alban. Exped. Pashtrik 1918 / 29.V-4.VI. / albanicus Bernh. Cotypus. / ex coll. Scheerpeltz / Dr M Bernhauer donavit / Cotypus Quedius albanicus Dr Bernhauer / Lectotypus ♂ Quedius albanicus Bernhauer, desig. V. Assing 2017 / Quedius illyricus Wendeler , det. V. Assing 2017" ( NHMW). Paralectotypes: 1♂, 4♀♀: "Alban. Exp. '18., Pashtrik 4-14.7. / Cotypus" ( NHMW) ; 14♀♀: "Alban. Exped. Pashtrik 1918, 29.V- 4.VI. / Cotypus" ( NHMW) ; 1♂: "Alban. Exped., Gjalica Ljums, 17.- 26.VI.'18 / Quedius v. albanicus Brnh. n. v. / Cotypus / Q. (Sauridus) albaniculus [sic] Coiff., H. Coiffait det. 1967" ( NHMW) .

Q. schipkanus: Lectotype ♂, present designation [dissected prior to present study]: " ♂ / Usanna- Hütte, Schiptschenska-Plan., Schipka-Balkan / 1400-1800 m., F. Schubert leg., Sommer 1925 / Usanna / ex coll. Scheerpeltz / Typus Quedius schipkanus O. Scheerpeltz / Lectotypus ♂ Quedius schipkanus Scheerpeltz, desig. V. Assing 2017 / Quedius illyricus Wendeler , det. V. Assing 2017" ( NHMW) . Paralectotypes: 1♀: same labels as lectotype ( NHMW) ; 2♂♂, 2♀♀: "Rosalito-polje,

Schipka-Balkan, ca. 1900 m / Juli 1935, leg. Ing. K. Mandl / ex coll. Scheerpeltz / Cotypus Quedius

schipkanus O. Scheerpeltz / Quedius illyricus Erichson , det. V. Assing 2017" (NHMW).

A d d i t i o n a l m a t e r i a l e x a m i n e d: Serbia: 4♂♂, 6♀♀, Stara Planina , Babin Zub, 22.VIII.2009, leg. Stevanović (cAss) ; 1♂, 1♀, Bukovic Planina , Kozji GRB, 700 m, 17.VII.2009, leg. Stevanović (cAss) ; 2♂♂, Tara Planina , Zaovine , 920 m, 28.X.2008, leg. Stevanović (cAss). Bosnia-Herzegovina: 1♂, 12 km S Kladanj, Konjuh planina, 950 m, beech and fir forest, 16.X.2005, leg. Hlaváč (cAss). Greece: 3♀♀, Flórina, 15 km WNW Flórina, Oros Varnous , 40°49'N, 21°15'E, 2000 m, 23.V.2005, leg. Assing (cAss) ; 4♂♂, 2♀♀, Flórina , 15 km WNW Flórina, Oros Varnous , 40°48'N, 21°15'E, 2010 m, 23.V.2005, leg. Assing (cAss) ; 1♂, Flórina, Pisodoria , 30.VII.2009, leg. Eifler (cAss); 2♀♀, Makhedonía , Kavála , West Rodopi , Skaloti env., Karandere Forest, Elatia old forest village, 1500-1600 m, 13-14.VI.2002, leg. Brachat (cAss) .

C o m m e n t: The original description of Q. albanicus is based on numerous ("in Anzahl") syntypes collected "am Pashtrik und Gjalica in Albanien " ( BERNHAUER 1926). The name is a junior primary homonym of Q. albanicus BERNHAUER, 1914 and was subsequently replaced with the nomen novum Q. illyricus by WENDELER (1928). COIFFAIT (1967) illustrated the aedeagus of a type specimen. As many as 28 syntypes, 27 of them females, were located in the collections of the NHMW. The sole male available is designated as the lectotype. Aside from Albania, the species was later reported from South Bulgaria ( KORGE 1969).

According to the original description, which is based on a unique specimen from " Albanien: Ljuboten", Q. paganettii is similar to Q. pyrenaeus BRISOUT DE BARNEVILLE, 1863, but distinguished by larger size ( BERNHAUER 1936). The species was subsequently reported also from Bosnia-Herzegovina and Macedonia by KORGE (1964). The description does not indicate any characters suggesting that it should be distinct from Q. illyricus .

Quedius schipkanus was described based on " 2♂♂ (1 Typus, 1 Cotypus) und 5♀♀ (1 Typus, 4 Cotypen) von Usanna, Schiptschenska-Planina, Schipka-Balkan", " 5♂♂, 13♀♀ vom Rosalito-polje, Schipka-Balkan", " 4♀♀ vom Massiv des Jumruktschal, Zentral- Balkan", " 1♀ von der Schiptschenska-Planina, Schipka-Balkan", and one ♂ and one ♀ from "Jumruktschal, Zentral-Balkan" ( SCHEERPELTZ 1937). Since a holotype is not specified, all the type specimens have syntype status. The male labelled by Scheerpeltz as "Typus" is designated as the lectotype. According to SCHEERPELTZ (1937), Q. schipkanus is closely allied to Q. umbrinus . A revision of the type material, however,

revealed that it is identical to Q. illyricus . COIFFAIT (1967) described Q. pseudopyrenaeus based on a male holotype from "Île de

Krk (Veglia)" and four female paratypes from Bosnia-Herzegovina. The type material deposited in the Coiffait collection is currently not accessible for taxonomic revision (see introduction). However, based on the original description, including the illustrations of the aedeagus, there is no doubt that the type material is conspecific with Q. illyricus . COIFFAIT (1978a) reported Q. pseudopyrenaeus also from Bosnia-Herzegovina and Serbia. KORGE (1969) already suspected the synonymy of Q. pseudopyrenaeus and Q. paganettii with Q. illyricus without formally synonymizing them, but was doubtful regarding the identities of Q. schipkanus and Q. bulgaricus.

The currently known distribution of Q. illyricus ranges from Croatia across Bosnia- Herzegovina, Serbia, Macedonia, and Albania to North Greece (first record!) and Bulgaria ( Map 2 ).

Figures

Map 2: Distribution of Quedius illyricus in the Balkans, based on revised (black circles) and literature records (white circles).