Raveniola zyuzini, Zonstein, 2024

Zonstein, Sergei L., 2024, A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia, European Journal of Taxonomy 967, pp. 1-185 : 57-59

publication ID	https://doi.org/ 10.5852/ejt.2024.967.2699
publication LSID	lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738
DOI	https://doi.org/10.5281/zenodo.14034712
persistent identifier	https://treatment.plazi.org/id/B61B579D-3D60-4EF1-A77D-5F7285F6356D
taxon LSID	lsid:zoobank.org:act:B61B579D-3D60-4EF1-A77D-5F7285F6356D
treatment provided by	Plazi
scientific name	Raveniola zyuzini
status	sp. nov.

Treatment

Raveniola zyuzini sp. nov.

urn:lsid:zoobank.org:act:B61B579D-3D60-4EF1-A77D-5F7285F6356D

Figs 18 View Figs 10–18 , 53 View Figs 45–53 , 73–74 View Figs 69–75 , 100 View Figs 100–108 , 127 View Figs 127–135 , 155 View Figs 148–159 , 185 View Figs 184–195 , 219 View Figs 211–219 , 246 View Figs 238–246 , 277 View Figs 274–281 , 337–339 View Figs 334–348 , 368–369 View Figs 364–378 , 436–438 View Figs 429–438 , 475–477 View Figs 475–486 , 532–533 View Figs 522–536 , 595–596 View Figs 593–601 , 711–714, 755

Diagnosis

Raveniola zyuzini sp. nov. differs from other members of the same species group by having a darker brownish orange carapace and legs (vs considerably paler brownish or yellowish orange ones – Figs 18 View Figs 10–18 , 53 View Figs 45–53 cf. Figs 15–17 View Figs 10–18 , 50–52 View Figs 45–53 ). Unlike males of R. diluta sp. nov. and R. fedotovi , possessing a raised embolic keel, males of R. zyuzini either entirely lack this structure or possess only a low vestige of the keel; the new species differs from R. pallens sp. nov. in having a longer proximal section of the embolus ( Figs 236–238 View Figs 229–237 View Figs 238–246 cf. Figs 234, 235 View Figs 229–237 ). Females of R. zyuzini are distinguishable due to their weakly sclerotized spermathecal trunks, carrying fusiform outer branches (vs differently conformed spermathecae in other species of the group; Figs 532–533 View Figs 522–536 cf. Figs 526–531 View Figs 522–536 ).

Etymology

The specific epithet is given in honour and memory of Dr. Alexei Zyuzin (1951–2021), noting his significant contribution to the taxonomical and faunistic study of Central Asian spiders.

Material examined

Holotype

UZBEKISTAN • ♂; Babatag Mts , environs of Chorroha (Chorraga) Pass, 10.4 km ENE of Mt Zarkassa; 38°04.0′ N, 68°13.5′ E; 1400 m a.s.l.; 15 Apr. 1990; S. Zonstein leg; SMNH. GoogleMaps

Paratypes (8 ♂♂, 4 ♀♀)

UZBEKISTAN • 1 ♂; same collection data as for holotype; SMNH GoogleMaps • 2 ♂♂, 1 ♀; same collection data as for preceding; 12 Apr. 1989; SMNH GoogleMaps • 1 ♂; same collection data as for preceding, northern slope of Mt Zarkassa ; 38°02.0′ N, 68°11.7′ E; 1900 m a.s.l.; 20 Apr. 2019; S. Zonstein leg.; SMNH GoogleMaps • 4 ♂♂, 3 ♀♀; same collection data as for preceding, eastern slope of Mt Zarkassa ; 1300–1800 m a.s.l.; 26 Apr. 1994; S.V. Ovchinnikov leg.; SMNH GoogleMaps .

Description

Male (holotype)

HABITUS. See Fig. 18. View Figs 10–18

MEASUREMENTS. TBL 12.20, CL 5.14, CW 4.32, LL 0.41, LW 0.86, SL 2.57, SW 2.26.

COLOUR. Carapace, as well as most palp and leg segments medium brownish orange; leg I slightly darker than other legs; eye tubercle blackish brown; chelicerae medium reddish brown; sternum, labium, maxillae, metatarsi and tarsi II–IV light yellowish orange; abdomen and spinnerets very pale yellowish brown, dorsal abdominal pattern indiscernible.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 100 View Figs 100–108 . Clypeus and eye group as in Fig. 155 View Figs 148–159 . Eye diameters and interdistances: AME 0.12(0.18), ALE 0.20, PLE 0.14, PME 0.13; AME–AME 0.12(0.06), ALE–AME 0.08(0.05), ALE–PLE 0.05, PLE–PME 0.03, PME–PME 0.31. Anterior cheliceral edge with unmodified setae; rastellum not developed. Each cheliceral furrow with 10 promarginal teeth and 3 mesobasal denticles. MIT indiscernible. Sternum, labium and maxillae as shown in Fig. 219 View Figs 211–219 . Maxillae with 8 cuspules each.

LEGS. Tibia and metatarsus I as in Fig. 277 View Figs 274–281 . Scopula: entire and distal on metatarsi I–II; entire on tarsus I; narrowly divided by setae on tarsus II; absent on tarsi III and IV. Trichobothria: 2 rows of 8–10 each on tibiae, 16–18 on metatarsi, 13–14 on tarsi, 10 on cymbium. PTC I–II and III–IV with 8–9 and 9–10 teeth on each margin.

SPINATION. Palp: femur d5, pd2, rd1; patella pd1; tibia d3, p3, r3, v7; cymbium d5(4). Leg I: femur d4, pd3, rd3; patella p1; tibia p2, pv2, r1(0), rv2+2M; metatarsus v1. Leg II: femur d4, pd3, rd 2(1); patella p1; tibia p3, v7; metatarsus p1, v6. Leg III: femur d4, pd3, rd3; patella p2, r2; tibia d2, p3, r3, v6; metatarsus d3, p3, r4, v7. Leg IV: femur d4, pd3, rd3(2); patella p1, r2; tibia d2, p3, r3, v7; metatarsus d2, p3, r3, v9. Tarsi I–IV aspinose.

PALP. Tibia, cymbium and copulatory bulb as shown in Fig. 368 View Figs 364–378 . Embolus with long, slightly tapering proximal section, provided with low rounded keel, and with short curved apex ( Fig. 436 View Figs 429–438 ).

SPINNERETS. See Fig. 595 View Figs 593–601 . PLS: maximal diameter 0.34; length of basal, medial and apical segments 0.78, 0.39, 0.34, respectively; total length 1.51; apical segment triangular.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	2.73 (2.42)	1.59 (1.57)	2.29 (1.85)	–	0.83 (1.63)	7.44 (7.47)
Leg I	4.51 (3.46)	2.60 (2.21)	3.57 (2.74)	3.62 (2.30)	2.22 (1.57)	16.52 (12.28)
Leg II	4.22 (3.28)	2.37 (1.98)	3.37 (2.40)	3.34 (2.28)	2.10 (1.55)	15.40 (11.49)
Leg III	3.63 (2.92)	1.96 (1.61)	2.78 (2.16)	3.75 (2.97)	2.09 (1.57)	14.21 (11.23)
Leg IV	4.72 (3.82)	2.24 (1.99)	3.71 (3.31)	5.38 (4.27)	2.43 (1.57)	18.29 (14.96)

Female (paratype)

HABITUS. See Fig. 53. View Figs 45–53

MEASUREMENTS. TBL 13.00, CL 4.42, CW 3.73, LL 0.36, LW 0.87, SL 2.12, SW 2.01.

COLOUR. In general as in male, but legs more uniformly coloured, without difference between legs I and II–IV.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 127 View Figs 127–135 . Clypeus and eye group as in Fig. 185 View Figs 184–195 . Eye diameters and interdistances: AME 0.10(0.14), ALE 0.21, PLE 0.12, PME 0.11; AME–AME 0.12(0.08), ALE–AME 0.09(0.07), ALE–PLE 0.05, PLE–PME 0.03, PME–PME 0.33. Cheliceral rastellum absent. Each cheliceral furrow with 9 promarginal teeth and 3–4 mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 246 View Figs 238–246 . Maxillae with 10–11 cuspules each.

LEGS. Scopula: entire and distal on metatarsi I–II; entire on palpal tarsus and tarsus I; narrowly divided by setae on tarsus II; absent on tarsi III and IV. Trichobothria: 2 rows of 9–10 each on tibiae, 13–14 on metatarsi, 11–13 on tarsi, 9–10 on palpal tarsus. Palpal claw with 4 promarginal teeth. PTC I–II and III–IV with 7–8 and 5–8 teeth on each margin, respectively.

SPINATION. Palp: femur d3, pd1; patella p1; tibia v7; tarsus v2(1). Leg I: femur d4, pd1; tibia v7; metatarsus v6(5). Leg II: femur d4, pd1; tibia p2, v7; metatarsus p1, v6. Leg III: femur d4, pd3, rd2; patella p1; tibia d1, p2, r2, v7; metatarsus d2, p3, r3, v7. Leg IV: femur d4, pd2(1), rd1; patella p1, r1; tibia d1, p3, r3, v7; metatarsus d1, p4, r4, v7. Patellae I–II and tarsi I–IV aspinose.

SPERMATHECAE. Each of paired spermathecae with wide cone-shaped trunk and short fusiform outer branch diverging from medium part of spermathecal trunk ( Figs 532–533 View Figs 522–536 ).

SPINNERETS. See Fig. 596 View Figs 593–601 . PMS: absent. PLS: maximal diameter 0.59; length of basal, medial and apical segments 0.68, 0.52, 0.47; total length 1.67; apical segment triangular.

Variation

Carapace length in males (n=7) varies from 3.80 to 5.19, in females (n =4) from 3.44 to 5.32. Within the type series, a few insignificant variations in the habitus and colouration are shown in Figs 73–74 View Figs 69–75 . For some structural details of the tarsal organ and trichobothria, see Figs 377–379 View Figs 364–378 View Figs 379–388 . The most common conformation of the copulatory bulb, lacking the keel, is similar to that shown in Figs 475–477 View Figs 475–486 . The less distributed variant of the bulb, provided with a rudimentary keel, is shown in Figs 369 View Figs 364–378 , 437–438 View Figs 429–438 .

Ecology

All spiders were found under stones in open park woodland composed of low deciduous trees ( Acer spp. , Crataegus spp. , etc.) at 1300–1600 m a.s.l. and in sparse mixed forest biotopes dominated by Juniperus seravschanica at 1600–1900 m a.s.l. See Figs 711–714 View Figs 707–714 .

Distribution

Known only from the type locality. See Fig. 755 View Figs 751–760 .