Reicheiodes (Himalayodes) subcirculatus, Michael Balkenohl & Joachim Schmidt, 2016

Michael Balkenohl & Joachim Schmidt, 2016, Reicheiodes GANGLBAUER, 1891 from Nepal: Description of a new species, and supplemental iconography of the Himalayan species (Insecta: Coleoptera: Carabidae: Dyschiriini), Contributions to Natural History 32, pp. 1-13 : 5-9

Reicheiodes (Himalayodes) subcirculatus sp. nov.

Fig. 1 .

Type material: Holotype, ♂, with label data “ NEPAL east Mangalbare dist. Terhatum 2.6.– 9.6.2013 lgt. E. Kučera ”, “Coll. P. Bulirsch” ( CBP) .

Diagnosis. A species with long-oval elytra, more regularly globose pronotum with less rounded elongation towards base, and moderately convex eyes with small genae. Chaetotaxy of the elytron is as follows: 1 basal, 1 subhumeral, 2 umbilical, 1 praeapical, and 1 dorsal setigerous punctures. Distinguished from the most similar species R. concameratus by the more slender total appearance, the absence of the second dorsal setigerous puncture, the shape of the pronotum with subcircular outline, the shape of the elytra with its maximum width at middle, the less convex elytra (lateral view), the more impressed punctures of the elytral striae which are in addition more distinctly impressed up to the apex, and the different shape of the endophallus. Another similar species is R. ellipsoideus which is distinguished by the much finer lateral channel of the pronotum, the presence of two dorsal setigerous punctures on the elytron, and the different shape of the endophallus. The geographically close species R. convexipennis is rather easily distinguishable from the new species because it exhibits three subhumeral setigerous punctures, and the pronotum is strikingly depressed with the lateral margin distinct at the anterior and posterior setigerous punctures only.

Measurements: Length 2.6 mm, width 0.96 mm, ratio length/width of pronotum 0.95, ratio length/width of elytra 1.45.

Colour: Head, pronotum, and dorsal surface dark reddish brown. Clypeus, vault of supraantennal plates, and clypeal field medium brown. Mandibles medium brown with carinae and apices darkened, mandibular and maxillary palpi medium brown with tip of apex light beige, femora and tibiae medium brown, tarsomeres medium brown, seven apical antennomeres dark brown, antennomeres 1–3 medium brown, fourth antennomere darkened apically. Ventral surface middle to dark brown.

Head: A third smaller than pronotum. Clypeus and lateral tooth distinctly margined. Clypeus straight, lateral tooth projecting, obtuse at tip, divided from supraantennal plates by obtuse but distinct notches; clypeal field nearly square, convex, smooth, separated from frons by deep straight transverse furrow; frons moderately convex, smooth; supraantennal plates convex, with carina at top of vault. Frontal furrows deep, broad, diverging anteriorly and posteriorly of transverse furrow. Eyes regularly convex; facets distinct, convex; genae enclosing eyes posteriorly by less than 10%. Antennae moderately long, extending beyond posterior setigerous puncture of pronotum, scapus with a single apical seta situated dorsally, antennomeres 5–10 moniliform. Labrum 7-setose, with indistinct isodiametric reticulation, nearly smooth. Mandibles moderately slender, arcuate apically. Terminal segments of maxillary and labial palpi securiform, both robust.

Pronotum: Outline subglobose, wider than long, maximum width at middle, in lateral view moderately convex. Lateral border subcircular in appearance, equally convex in middle part, narrowed with slightly rounded elongation to base. Reflexed lateral margin distinct, reaching from rounded anterior angles up to posterior setigerous puncture, extended over puncture by half of distance to base, joining anterior transverse line. Lateral channel moderately broad and deep. Median line sharp, distinct, deeper at base, joining anterior transverse line; anterior transverse line complete, developed as moderate line; surface shiny, with rough wrinkles laterally, with few subtle pierced punctures, flange of moderate size.

Elytron: Convex on disc, moderately to slightly convex in anterior part, more distinctly convex at base (lateral view). Outline long-oval, maximum width at middle, margined from pedunculus to apex; no humeral angle traceable; lateral channel moderately broad from level of humerus to apex, fine at base; reflexed margin distinct. Basal granula absent; basal setigerous puncture distinct, situated in projected extension of first stria. One subhumeral, two umbilical, one praeapical setigerous puncture(s). Parascutellar stria fine, situated at base. First stria reaching basal setigerous puncture, fifth reaching base, others ending basally at declivity, first one joining lateral channel at apex; inner five striae impressed, all formed by row of impressed punctures, becoming less impressed towards apex and laterally. All intervals moderately convex. Third interval with one (anterior) setigerous puncture, approaching second stria.

Hind wings: Completely atrophied.

Ventral surface: Proepisternum with very fine wrinkles, almost smooth. Abdominal sternites nearly smooth, terminal segment with indistinct and very fine rugae-like reticulation, two apical setigerous punctures widely distant.

Protibia: Lateral upper spine curved ventro-laterally. Movable spur smaller than spine, gently curved. Praeapical lateral denticle strong, sharp, second one much smaller.

Male genitalia ( Figs 7, 8 ): Median lobe moderately sclerotized, distinctly and angle like arcuate in middle part, slightly flattened in apical half, apex formed by asymmetric rounded spatula, dorsally and ventrally with very few fine pili in basal half. Oroficium medium sized, closing lips less sclerotized. Endophallus with numerous wrinkles, with three small spines in basal part (visible at 500 times by optimized condenser with narrow-band filter green 546 nm). Parameres asymmetrically, length of the ventral one less than a third of the dorsal one, ventral one slightly twisted, dorsal one moderately twisted, both with a short seta at apex.

Female genitalia: Unknown.

Etymology. The name refers to the subcircular appearance of the pronotum.

Distribution ( Fig. 12 ). Up to today only known from the western slope of the Singalila mountain range in East Nepal. Based on the label data, the single specimen was found east of the village Mangalbare, which is situated in the Ilam District, but not in the Terhatum District, as erroneously stated on the locality label.

Habitat. Unknown.

New records

Figures

Fig. 12: Distribution of the species of the Reicheiodes subgenus Himalayodes in Nepal (recorded localities of all species plotted).

Figs 1 – 4: Habitus. 1: Reicheiodes (Himalayodes) subcirculatus sp. nov., holotype, ♂. 2: R. convexipennis BALKENOHL, 1994, paratype, ♂. 3: R. loebli BALKENOHL, 1994, holotype, ♂. 4: R. ellipsoideus BALKENOHL, 1995, holotype, ♀.

Figs 7 – 8: Reicheiodes (Himalayodes) subcirculatus sp. nov., holotype; 7: Aedeagus with median lobe and parameres, ventral view. 8: Apical part of median lobe, ventral view.