Rhinocladius culicinus Edwards

Saether, O. A. & Andersen, T., 2003, Redescription of Rhinocladius Edwards (Diptera: Chironomidae: Orthocladiinae), Zootaxa 217, pp. 1-20 : 4-6

Rhinocladius culicinus Edwards ( Figs 1­8 )

Rhinocladius culicinus Edwards 1931: 271 .

Material examined: CHILE: Terr. Rio Negro., L. Correntoso, holotype Ψ: 18­25 xi 1926, F. & M. Edwards ( BMNH 1927­63); Llanquihue Prov., Casa Pangue, 3 Ψ paratypes, 4­10 xii 1926, F. & M. Edwards ( BMNH 1927­63). (According to Edwards, 1931, however, one of the paratypes is supposed to be from L. Correntoso.

Diagnostic characters: The imagines differ from the only other known species, R.. longirostris , by having labellae 0.50­0.56 times as long as the wing, several setae on postcubitus, and fewer prealars.

Female imago (n=3­4 except when otherwise stated)

Total length 2.60­2.98 mm. Wing length 1.70­2.10, 1.88 mm long. Total length/wing length 1.51­1.60. Wing length/length of profemur 2.32­2.43. Thorax, head and legs dark brown; abdomen dark with posterior 1/4 more pale.

Head ( Fig. 2 ). AR 0.42­0.44. Length of flagellomeres (in m) as: 68­83, 77; 71­94, 84; 71­105; 71­98; 64­83; 154­188. Temporal setae 11­13, 12; consisting of 7­9, 8 inner verticals; 2­3, 2 outer verticals; and 2 postorbitals. Clypeus with 12­21 setae . Tentorium ( Fig. 1 ) 173­210, 189 m long: 19­24, 21 m wide; stipes 83­131 m long, 23­26 m wide. Palpomere as in R. longirostris ( Fig. 20 ), lengths (in m): 11; 23­34, 28; 60­79, 71; 26­30, 29; 21­30, 25. Third palpomere with 16­24 (2) spatulate sensilla clavata, fourth palpomere with 3­4 (2) spatulate sensilla clavata. Labellae 869­1106 m long, 41­58 m wide, 0.50­0.56 times as long as wing.

Thorax ( Fig. 3 ). Antepronotum with 4 lateral setae . Dorsocentrals 17­24, 21; acrostichals 14­18 (2); prealars 7­8, 8. Scutellum with 12­14, 14 setae .

Wing. VR 1.36­1.47, 1.42. Costal extension 143­169, 157 m long. R with 22­40, 32 setae ; R1 with 33­55, 41; R4+5 with 49­94, 67; M with 16­25, 20; RM with 1­3,2; M1+2 with 70­93, 83; M3+4 with 36­58, 44; Cu with 30­60, 41; Cu1 with 19­32, 24; postcubitus with 8­22, 14; An with 27­46, 35; and costal extension with 16­20 non­marginal setae . Cell m with 1­12, 6 setae ; r4+5, m1+2, and cu plus an each with about 250­300 setae ; m3+4 with about 75­95 setae . Squama with 7­10, 8 setae

Legs. Spur of front tibia 34 m (2) long; spurs of mid tibia 30 (1) and 23­26 m long; spurs of hind tibia 83­94 and 23­30 m (2) long. Width at apex of front to hind tibiae (in m) as: 41­49; 41­49, 45; 53­56. Comb of 14­15 setae , shortest seta 34­38 m long, longest seta 75­94 m long. Lengths (in m, n= 2­4) and proportions of legs:

Abdomen. Number of setae on tergites I­VIII as: 44­50; 36­56; 46­55; 40­55; 36­58; 37­51, 44; 18­43, 32; 18­27, 23. Number of setae on sternites I­VIII as: 0; 2; 11­14; 21­25; 25­33; 30­50, 37; 25­41, 35; 34­41, 39 median and 6­8, 5 lateral.

Genitalia ( Figs 4­8 ). Tergite IX with 22­32, 27 setae . Gonocoxite with 6 setae including 1 long. Notum 83­90, 86 m long. Cercus 83­105, 93 m long. Seminal capsule 83­90, 86 m long; with 25­30 m long neck; 60­79, 71 m wide.

Remarks

R. culicinus is very similar to R. longirostris described below differing significantly only in the shorter labellae and the presence of setae on postcubitus. It thus is not unlikely that the species redescribed below represents only a form of R. culicinus .

Figures

FIGURES 1 ­ 8. Rhinocladius culicinus Edwards, female imago, 1: cibarial pump, tentorium and stipes, 2: head, 3: thorax, 4: genitalia, ventral view, 5: apodeme lobe, 6: ventrolateral lobe, 7: dorsomesal lobe, 8: genitalia, dorsal view.

FIGURES 16 ­ 21. Rhinocladius longirostris Edwards, female imago, 16: head, 17: cibarial pump and stipes, 18: tentorium, 19: genitalia, ventral view, 20: palp, 21: genitalia, ventral view.