Scopaeus pulcher Frisch, 2022

Frisch, Johannes & Narakusumo, Raden Pramesa, 2022, Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species, Soil Organisms 95 (1), pp. 23-73 : 41-42

publication ID	https://doi.org/ 10.25674/so95iss1id311
DOI	https://doi.org/10.5281/zenodo.10724296
persistent identifier	https://treatment.plazi.org/id/03B7C77A-FFD2-FFE7-BCF5-FBDE0F6E0561
treatment provided by	Felipe
scientific name	Scopaeus pulcher Frisch
status	sp. nov.

Treatment

Scopaeus pulcher Frisch , spec. nov.

( Figs 13 View Figures 11–13 , 40 View Figures 35–41 , 83 –85 View Figures 80–85 , 130 View Figures 119–133 , 161, 162 View Figures 155–162 )

Type specimens: Indonesia, Jawa Barat: Holotype ♂, Sukabumi, Cikaniki: Cikaniki River ( Mt Halimun , Halimun-Salak NP), (06°44’46’’S, 106°32’25’’E), 1020 m, 17.9.2015, leg. Frisch ( MZB) GoogleMaps . Paratypes (145 specimens): 26 ♂, 14 ♀, same data as holotype ( MFNB, MZB) GoogleMaps ; 31 ♂, 27 ♀, Sukabumi, Cikaniki: Cikaniki River ( Mt Halimun , Halimun-Salak NP), (06°44’46’’S, 106°32’25’’E), 1020 m, 5.10.2015, leg. Frisch ( MFNB, MZB) GoogleMaps ; 2 ♂, 1 ♀, Sukabumi, Citalahab Central: Cikaniki River (06°44’22’’S, 106°31’55’’E), 1070 m, 12.5.2016, leg. Frisch ( MFNB, MZB) GoogleMaps ; 15 ♂, 9 ♀, Bogor, SW Pamijahan ( Cimuara Herang River ), (06°42’41’’S, 106°41’03’’E), 990 m, 3.10.2015, leg. Frisch ( MFNB, MZB) GoogleMaps ; 5 ♂, 12 ♀, Bogor, S Tenjolaja, Tapos : Ciampea River (06°41’36’’S, 106°42’24’’E), 1040 m, 2.10.2015, leg. Frisch ( MFNB, MZB) GoogleMaps ; 1 ♀, Bogor, Mt Salak: Sungai Ciapus , 800 m, 17.6.1994, leg. Schuh ( NHMW) ; 1 ♂, 1 ♀, Sukabumi, NNW Cidahu: Javana Spa (06°44’43’’S, 106°42’51’’E), 1150 m, 30.9.2015, leg. Frisch ( MFNB, MZB) GoogleMaps .

Description: Habitus and coloring as in Fig. 13 View Figures 11–13 . Head subcircular. Penultimate antennal segment elongate. Mesotibia moderately thickened. Setose punctation of forebody surface spacious, fine on head, absent on pronotum, very coarse in median fourth of elytral width, in proximal fourth of elytra with coarse, close punctures in narrow strip along suture; elytral punctures becoming considerably smaller laterad and extinguished in polished lateral third of elytra down to ventral margin; abdomen finely, densely punctate, matter than shiny forebody; microreticulation absent. Forebody setation long and raised with numerous macrosetae not much longer than fine pubescence. Coloration variable; body orange-brown; head somewhat darker than pronotum; elytra in proximal third darker orange-brown than head, followed by transverse, blackish-brown band extended towards shoulders laterally and light orange-brown posterior fifth; abdomen dark orange-brown with disc of segments blackish in variable extent; maxillary palpi with penultimate segment blackish; antennae with orange-brown scapus, blackish pedicellus and median segments, gradually lighter towards terminal segments; dark color components frequently reduced up to almost unicolorous, orange-brown specimens with somewhat lighter pronotum, darker elytra and light yellow-brown appendages. Total body length 3.3 – 4.0 mm; forebody length 2.0 – 2.2 mm.

Male: Abdominal sternite VII with unmodified posterior margin. Emargination of abdominal sternite VIII bisinuate with convex middle, occupying about posterior seventh of sternite length ( Fig. 130 View Figures 119–133 ). Aedeagus ( Figs 40 View Figures 35–41 , 83 – 85 View Figures 80–85 ) about 0.8 mm long with asymmetrical distal lobes; dextral apical lobe elongate with ventrally extended, convex apex ( Figs 40 View Figures 35–41 , 83 View Figures 80–85 ), slender in ventral and dorsal view ( Figs 84, 85 View Figures 80–85 ); sinistral apical lobe much wider, strongly extended dextrad distally and overlapping end of dextral apical lobe ( Figs 84, 85 View Figures 80–85 ); dorsal lobe with remarkable, sinistral enlargement stepwise narrowed proximad and projecting from apical lobes ventrally ( Figs 40 View Figures 35–41 , 83 View Figures 80–85 ); flagellum short, not projecting from apical lobes, bent sinistrad towards end ( Figs 40 View Figures 35–41 , 83, 84 View Figures 80–85 ); ventral lobe short, emarginate at apex with distad curved ends of unequal length ( Figs 40 View Figures 35–41 , 83, 84 View Figures 80–85 ); lateral lobes reduced, marked by group of setae ( Figs 84, 85 View Figures 80–85 ); median foramen small, asymmetrical, with transverse ridge with lateroproximally curved ends and proximad curved middle projecting ventrally ( Figs 40 View Figures 35–41 , 83 View Figures 80–85 ).

Female: Sperm pump as in Fig. 162 View Figures 155–162 ; bursal duct about twice as long as sperm pump; bursa with sclerotized end of specific shape ( Fig. 161 View Figures 155–162 ).

Phylogeny: Scopaeus pulcher is a representative of the S. gracilis species group ( Frisch et al. 2002: 39).

Distribution: The type specimens of Scopaeus pulcher originate from Mount Halimun and Mount Salak, Jawa Barat. A specimen from northern Sumatra (Lake Toba, leg. Schillhammer, NHMW) might point to a much wider distribution, but more specimens are necessary to judge if the slight genital differences are within the range of intraspecific variation.

Etymology: The epithet pulcher (adjective, Latin: ‘beautiful’) denotes the multicolored coloration of this nice, new species ( Fig. 13 View Figures 11–13 ) in contrast to the usually more uniformly colored majority of Scopaeus .