Slaterocoris sheridani Knight

Schwartz, Michael D., 2011, Revision And Phylogenetic Analysis Of The North American Genus Slaterocoris Wagner With New Synonymy, The Description Of Five New Species And A New Genus From Mexico, And A Review Of The Genus Scalponotatus Kelton (Heteroptera: Miridae: Orthotylinae), Bulletin of the American Museum of Natural History 2011 (354), pp. 1-290 : 223-229

Slaterocoris sheridani Knight

Figures 13 , 45A–C , 46 , 47 ; plates 3 , 4O; map 9

Strongylocoris robustus: Van Duzee, 1921: 136 (disc. in part).

Slaterocoris sheridani Knight, 1968: 92 , fig. 113 (orig. desc.); Kelton, 1968: 1125, fig. 8 (disc.); Knight, 1970: 255, fig. 2 , (disc.); Henry and Wheeler, 1988: 451 (catalog); Schuh, 1995: 199 (catalog).

Slaterocoris dakotae Knight, 1970: 243 , fig. 4 (orig. desc.); Henry and Wheeler, 1988: 449 (catalog); Schuh, 1995: 197 (catalog). NEW SYNONYMY.

Slaterocoris texanus Knight, 1970: 252 , fig. 39 (orig. desc.); Henry and Wheeler, 1988: 451 (catalog); Schuh, 1995: 199 (catalog). NEW SYNONYMY.

Slaterocoris basicornis Knight, 1970: 254 , fig. 6 (orig. desc.); Henry and Wheeler, 1988: 449 (catalog); Schuh, 1995: 197 (catalog). NEW SYNONYMY.

Slaterocoris custeri Knight, 1970: 254 , fig. 3 (orig. desc.); Henry and Wheeler, 1988: 449 (catalog); Schuh, 1995: 197 (catalog). NEW SYNONYMY.

Slaterocoris woodgatei Knight, 1970: 254 , fig. 5 (orig. desc.); Henry and Wheeler, 1988: 452 (catalog); Schuh, 1995: 200 (catalog). NEW SYNONYMY.

DIAGNOSIS: Distinguished from other Slaterocoris species by the following combination of characters: coxae and trochanters black, femora black except for pale yellowish orange apical regions, tibiae pale yellowish orange or brown, tarsomeres black (fig. 13); vestiture of sparse, moderately short, reclining, dark simple setae (pl. 4O); antennal segments I and II usually black, sometimes segment I pale or dusky yellow (pl. 3); right paramere with entire (fig. 43C, D) or bifurcate (fig. 43B, E–G) medial process; twisted or concave dorsal lobe of endosomal spicule somewhat broad and gradually attenuate (fig. 44C–H). The proceeding features will also distinguish S. sheridani from other flavipes -group species, as S. sparsus has entirely pale yellowish femora (fig. 13) and S. flavipes and S. longipennis have dense and pale vestiture. Slaterocoris sheridani could be confused with members of the breviatus group, but these latter species are practically glabrous and tarsomeres I and II are pale (fig. 13). From apache -group species, the more sharply contrasting pale apical and dark portions of the femora will help discriminate S. sheridani (fig. 13) from S. croceipes , which has variable width orangebrown medial regions between the black bases and pale yellow apices (fig. 12). Slaterocoris apache has similar femoral coloration to S. sheridani , but has suberect, longer, pale dorsal vestiture (pl. 4B). In the female the shorter average antennal segment II of S. sheridani (0.82, 0.78–0.93) will further distinguish it from these apache -group species with longer average segment II (1.05, 0.96–1.13 apache ; 1.02, 0.78–1.23 croceipes ).

REDESCRIPTION: Male: Moderately small, subovate; length 3.35–4.19, width 1.60–1.81 (pl. 3). COLORATION: Black; including antennal segments I and II, sometimes segment I and II mostly dusky pale yellowish to orange-brown basally; leg black, apical region of femur and tibia pale yellowish to orange-brown, distal region of femur without gradual tint progression from black to orange to yellow, color changes sharply from black to yellow (fig. 13). VESTITURE: Sparse, reclining, moderately short, brown to golden brown simple setae (pl. 4O). STRUCTURE: Body conformation roughly similar in both sexes; costal margin slightly curved; antennal segment II shorter than foretibia. MALE GENITALIA: As in species group except: Endosomal spicule: Dorsal lobe with narrow dorsal portion longer than wide basal portion; serration of basal portion sparse; apex of dorsal portion somewhat narrow (fig. 45A–C, arrow). Right paramere: Subapical region long with variable number of small spines and prominent and variable medial lobe, medial lobe either entire (fig. 46A–E, G, H arrow, L) or bifurcate (fig. 46F arrow, I–K).

Female: Moderately small, ovate; length 3.45–4.15, width 1.70–2.13 (pl. 3). COLORATION: As in the male, but antenna usually paler, antennal segment I extensively dusky pale basally, segment II widely pale medially, darker basally and distally; III and IV black; apices of femur with narrow orange-brown area between black basal and pale yellow apical regions. STRUCTURE: Costal margin arcuate. GENITALIA: As in species group except: Vestibulum, anteroventral margin of anterior wall: Sclerotized and incised medially, sclerotization reaching to lateral margins of bursa copulatrix (fig. 47B). Ventral labiate plate: Strongly produced, base moderately wide, triangular, apex cleft (fig. 47A). Second gonapophyses: Anterior medial surface slightly convex (fig. 47C). Interramal sclerite: Dorsomedial region convex anteriorly, and posteromedial portion with flat plate abutting ovipositor bulb (fig. 47D). Interramal lobe: Basal (dorsal) insertion with sclerotized attachment, dorsomedial margin with short process (fig. 47D).

HOSTS: Specimens taken from these Asteraceae : an unidentified genus and species (2 specimens, 1 collection event), an unidentified species of Artemisia (15 specimens, 1 collection event), Chrysothamnus (one or more unidentified species [30 specimens, 3 collection events] and C. viscidiflorus [4 specimens, 1 collection event]), Ericameria nauseosa (6 specimens, 2 collection events), and Gutierrezia sarothrae (15 specimens, 2 collection events). Large numbers of specimens were taken from G. sarothrae at localities in Utah (7 specimens) and Nebraska (8 specimens). Even more specimens were collected from unspecified species of Chrysothamnus at Arizona (29 specimens) and Utah (3 specimens) sites; however, it is not possible to determine whether these occurrences should be attributed to C. viscidiflorus (or a congener) or E. nauseosa , or both. Based on the number of specimens and collecting events it is probable, as in other western species of Slaterocoris , that both rabbitbrush species and at least one species of Artemisia are breeding hosts. Fraxinus cuspidata Torr. (Oleaceae) (15 specimens, 1 collection event) is probably not a breeding host, but may prove to be a factor to the biology of S. sheridani in the same manner that species of Lonicera and Symphoricarpos (Caprifoliaceae) are to S. pallidicornis . Only two specimens are recorded from non-asteraceous plants; these are considered to be sitting records.

DISTRIBUTION: The southern extent is in west Texas, reaching west to the Mogollon Rim of east and central Arizona, north through the Rocky Mountain cordillera of New Mexico to the front range of Colorado, extending to the northern Great Plains from northern Colorado, to Wyoming, and southern North Dakota. Sporadic localities occur in the intermountain west of Colorado and Utah with the westernmost sites in the isolated ranges of Nevada and southern Utah ( map 9 ).

COLLECTION SUMMARY: 335 specimens from 75 collection events, with the earliest records in April and May from the southern portion of the range in Arizona. Throughout the remaining distribution, collection dates cover June and July with August dates known from New Mexico and west Texas, as well as in the northern localities of South Dakota and Wyoming.

DISCUSSION: Knight (1968) described S. sheridani from Sheridan, Wyoming, based on two males taken in the same collection event. Subsequently, he similarly founded five additional species on a few specimens ( Knight,1970) as follows: S. basicornis —male holotype from Big Bend, Texas; S. custeri — male holotype and male paratype (same collection event) from Custer, South Dakota, and female paratype from nearby Sundance, Wyoming; S. dakotae —male holotype from Interior, South Dakota, S. texanus —male holotype from Big Bend, Texas; and S. woodgatei —male holotype and two male paratypes from Jemez Springs, New Mexico (same collection event). The six nominal species were purportedly distinguished from each other by the vestiture coloration, body size, coloration of antennal segment I, comparative length of antennal segment II to head width or foretibia, and structure of the right paramere. All the proposed discriminating features have proven inadequate after examining the holotypes and other specimens fitting the diagnoses of the six nominal species.

Knight (1968, 1970) variously described the vestiture for these taxa as pallid ( basicornis ), pale yellowish to golden ( custeri ), pale to golden ( texanus ), golden yellow ( sheridani ), golden ( woodgatei ), and brownish ( dakotae ). Kelton (1968) characterized the coloration of sheridani as brownish. Accurately determining the coloration of the dorsal vestiture is difficult because of its sparse distribution, short length, and the vagaries of illumination. I can find no clearcut geographically significant difference in the vestiture coloration; all are brown or golden brown. Body length in males spans 3.35 to 4.19 mm, encompassing the range of the holotypes of all the nominal species. Actually, the range (0.84) of the body length is smaller than in the related species S. sparsus (1.20), which is known only from southern California. The coloration of antennal segment I is variable across the distribution and within a collection event, ranging from pale yellow to black, with most specimens dusky yellowish brown; sometimes the dorsal aspect of the segment is paler than the ventral aspect. The length of antennal segment II is always shorter than the head width, but ranges from just shorter to almost one-third shorter.

The suggested structural differences in the right paramere (cf. Knight, 1970: 259, 265, figs. 2–6 , 39) of the nominal species is actually less than the variation in the number and conformation actually documented (cf. fig. 47A–L) during the course of this study from across the distribution of the nominal species from North Dakota to New Mexico and Colorado to Utah. The medial process is entire (fig. 46A–E, G, H, L) or bifurcate (fig. 46F, I, J, K) and the tubercle on the basal region is entire (figs. 46E, F) or serrate (fig. 46A–C, G–L). A male with a large basal process (cf. fig. 46D) is apparently aberrant as a another male from the same collection event has more typical morphology (fig. 46C). Compared with the right paramere the structural variation it the endosomal spicule is minor, limited to the intensity of minute serration on the base of the dorsal lobe (fig. 45A–C).

The type localities of three of the nominal species in the northern Great Plains of North Dakota and Wyoming are within 500 km of each other, whereas the two nominal species from Texas are from Big Bend National Park having been collected on consecutive days, with one species collected in north-central New Mexico. A female, in the CNC, from the same collection event as the holotype and paratype of S. woodgatei , has a Knight identification label of S. sheridani attached. Only minor differences are noted between the nominal species and all are well within the range incorporated in the reformed diagnosis of S. sheridani provided above. I am therefore treating S. basicornis , S. custeri , S. dakotae , S. texanus , and S. woodgatei as junior synonyms of S. sheridani .

Van Duzee (1921) mentioned that the CAS retained specimens of S. robustus from Boulder and Denver, Colorado. These are correctly identified as S. sheridani (see Other Specimens Examined).

HOLOTYPES: USA: New Mexico: Sandoval Co.: Jemez Springs [35.768616 N 106.691676 W, 1951 m], 07 Jun 1916 – 17 Jun 1916, unknown, 18 (AMNH_PBI 00134023) (USNM) ( S. woodgatei : junior synonym). South Dakota: Custer Co.: Custer [43.766676 N 103.598336 W], 19 Aug 1927, H.H. Knight, 18 (AMNH_PBI 00122243) (USNM) ( S. custeri : junior synonym). Jackson Co.: Interior [43.726946 N 101.983616 W], 28 Jun 1924, H.C. Severin, 18 (AMNH_PBI 00122245) (USNM) ( S. dakotae : junior synonym). Texas: Brewster Co.: Big Bend National Park, Green Gulch [29.33216 N 103.24186 W, 1200 m], 14 Aug 1969, Board and Hafernik, 18 (AMNH_PBI 00133387) (USNM) ( S. basicornis : junior synonym). South Rim Trail, Chisos Mountains, Big Bend National Park [29.229366 N 103.29546 W, 2118 m], 15 Aug 1969, Board and Hafernik, 18 (AMNH_PBI 00122266) (USNM) ( S. texanus : junior synonym). Wyoming: Sheridan Co.: Sheridan [44.797226 N 106.955566 W], 03 Aug 1927, H.H. Knight, 18 (AMNH_PBI 00134021) (USNM) ( S. sheridani : senior synonym).

OTHER SPECIMENS EXAMINED: USA: Arizona: Gila Co. : 2 mi W of Miami, 33.399176 N 110.902696 W, 1158 m, 04 May 1979, J.D. Pinto, E.M. Fisher, 18 (00119797) (UCR); 04 May 1979, J.D. Pinto, E.M. Fisher, 18 (00119891) (UCR). 12 mi W of Payson, 34.230836 N 111.534266 W, 975 m, 01 May 1981, D.A. and J.T. Polhemus, 78 (00116717, 00117259–00117264), 6♀ (00117207, 00117254– 00117258) (JTP). 15 mi N of Globe on Ariz. 88, 33.605326 N 111.005316 W, 914 m, 28 Apr 1981, D.A. and J.T. Polhemus, 18 (00116716) (JTP). Globe, 33.23396 N 110.47116 W, 1074 m, 09 May 1931, D.K. Duncan, Asteraceae , 18 (00242642), 1♀ (00242643) (CAS), 58 (00112158, 00112160–00112163), 9♀ (00112159, 00112164–00112171) (CNC), 1♀ (00164892) (USNM). Mazatzal Wild[erness Reserve] along E Verde River, 34.286396 N 111.664176 W, 30 Apr 1981, D.A. and J.T. Polhemus, 68 (00116715, 00117265–00117269), 3♀ (00117270–00117272) (JTP). Rye, 34.109446 N 111.353066 W, 1067 m, 01 May 1981, D.A. and J.T. Polhemus, 198 (00116718, 00117197, 00117285–00117301), 11♀ (00117198, 00117275– 00117284) (JTP). base of Pinal Mountains, 33.2896 N 110.8686 W, 1219 m, 30 May 1930, Parker and Duncan, Prosopis sp. (Fabaceae) , 2♀ (00164890–00164891) (USNM). Graham Co.: 36 mi SE of Globe on I-70, 33.252396 N 110.282346 W, 853 m, 28 Jun 1981, D.A. and J.T. Polhemus, 1♀ (00116985) (AMNH), 1♀ (00116759), 118 (00116708, 00117330–00117339), 23♀ (00117199, 00117308– 00117329) (JTP). Stockton Pass, Pinaleno Mountains, 32.640836 N 109.843066 W, 1631 m, 01 Jun 1983 – 02 Jun 1983, R.T. Schuh, G.M. Stonedahl, Fraxinus cuspidata Torr. (Oleaceae) , 108 (00107902, 00169995–00170003), 6♀ (00108420, 00108854, 00170004–00170007) (AMNH). Maricopa Co. : Fish Creek, Tonto National Forest, 33.53526 N 111.3066 W, 671 m, 09 May 1918 – 10 May 1918, J.C. Bradley, 68 (00116193 [cat Cornell Univ. Lot 542 Sub 130]–00116198 [cat Cornell Univ. Lot 542 Sub 130]), 13♀ (00116199 [cat Cornell Univ. Lot 542 Sub 130]–00116211 [cat Cornell Univ. Lot 542 Sub 130]) (CUIC). Salt River Canyon at Apache Lake, 33.558116 N 111.531536 W, 610 m, 28 Apr 1981, D.A. and J.T. Polhemus, 1♀ (00116832) (JTP). Sunflower, 33.86426 N 111.467636 W, 02 Jun 1981, J.T. Polhemus, 1♀ (00116831) (JTP); 23 May 1982, D.A. and J.T. Polhemus, Light Trap, 18 (00117273), 1♀ (00117274), 28 (00116726, 00116820) (JTP). nr Fish Creek on Rt 88, 33.5256 N 111.30716 W, 800 m, 29 Apr 1981, D.A. and J.T. Polhemus, 18 (00116731) (JTP). Navajo Co.: 15–20 mi SW of Show Low, 34.049726 N 110.275926 W, 1707 m, 30 May 1983, Schuh, Stonedahl, and Massie, Chrysothamnus sp. , 128 (00107903, 00108145, 00170008–00170017), 8♀ (00108962, 00170018– 00170024) (AMNH). Yavapai Co.: 1 mi E of Stoneman Lake, interchange on I-70, T16E R7E Sec19, 34.764466 N 111.656546 W, 15 Jun 1983, R.T. Schuh and M.D. Schwartz, Chrysothamnus sp. , 28 (00107901, 00107914), 6♀ (00108414–00108419) (AMNH), 1♀ (00119892) (UCR). 1 mi N of I-17 on Rt 179, T15N R6E Sec 18, 34.694236 N 111.752386 W, 1158 m, 15 Jun 1983, R.T. Schuh and M.D. Schwartz, 1♀ (00108413) (AMNH). 5 mi E of Camp Verde, 34.53736 N 111.81936 W, 975 m, 30 Apr 1981, J.T. Polhemus, 1♀ (00116833) (JTP). 5 mi S of Clarkdale, 34.7366 N 112.06616 W, 1158 m, 30 Apr 1981, D.A. and J.T. Polhemus, 38 (00116719, 00117302–00117303), 4♀ (00117304– 00117307) (JTP). Yarnell, Weaver Mountains, 34.215836 N 112.777786 W, 22 May 1937, L.K. Gloyd, 18 (00127469), 1♀ (00127470) (UMMC); 23 May 1937, L.K. Gloyd, 18 (00127471) (UMMC); 24 May 1937, L.K. Gloyd, 18 (00127472) (UMMC); 27 May 1937, L.K. Gloyd, 18 (00127466), 2♀ (00127467–00127468) (UMMC). Colorado: Arapahoe Co.: Cherry Creek Reservoir, 39.61346 N 104.8226 W, 1707 m, 21 Jun 1980, D.A. Polhemus, 1♀ (00116835) (JTP). Englewood, 39.647786 N 104.987226 W, 11 Jun 1978, D.A. and J.T. Polhemus, 1♀ (00116761), 48 (00116725, 00117235–00117237), 8♀ (00117200, 00117238–00117244) (JTP). Boulder Co.: Boulder, 40.0156 N 105.276 W, 1676 m, 21 Jul 1903, E.P. Van Duzee, 3♀ (00242644–00242646) (CAS). Denver Co.: Denver, 39.739176 N 104.984176 W, 12 Jul 1903, E.P. Van Duzee, 1♀ (00242725) (CAS). Douglas Co.: Roxborough Park Road near Chatfield State Park, 39.473896 N 105.084726 W, 1707 m, 04 Jun 1981, D.A. Polhemus, 18 (00116834) (JTP). nr Waterton, Roxborough Road, 39.493616 N 105.088066 W, 1707 m, 04 Jun 1981, D.A. Polhemus, 1♀ (00117246) (JTP); 22 Jun 1981, D.A. Polhemus, 1♀ (00117245) (JTP). Huerfano Co.: 5 mi W of Walsenburg, 37.59816 N 104.85816 W, 1981 m, 06 Jul 1982, D.A. and J.T. Polhemus, 1♀ (00116841) (JTP). Jefferson Co.: Golden, 39.755566 N 105.220566 W, 1829 m, 21 Jun 1961, W.R.M. Mason, 18 (00112156), 1♀ (00112156) (CNC). Red Rocks Park, 39.669726 N 105.202786 W, 23 Jun 1982, D.A. Polhemus, 18 (00116760) (JTP); 21 Jun 1988, D.A. Polhemus, 18 (00116838), 1♀ (00116839) (JTP). Montrose Co.: 18 mi SE of Naturita, 38.034326 N 108.334446 W, 08 Jul 1980, J.T. and D.A. Polhemus, 18 (00116840) (JTP). Pueblo Co.: 12 mi W of Pueblo off Hwy 96, 38.254446 N 104.829526 W, 15 Jun 1980, J.T. and D.A. Polhemus, 1♀ (00116986) (AMNH), 38 (00116710, 00116836–00116837), 10♀ (00116842–00116844, 00117247–00117253) (JTP). Weld Co.: 17.5 mi N of Nunn, Pawnee National Grassland, 40.95666 N 104.780286 W, 08 Jul 1973 – 09 Jul 1973, J.A. Slater and R. Baranowski, 18 (00107917), 2♀ (00108411– 00108412) (AMNH). Nebraska: Sioux Co.: 13 mi N of Mitchell on Rt 29, 42.127086 N 103.819816 W, 1387 m, 17 Jun 1998, A.G. Wheeler, Jr., Gutierrezia sarothrae , 58 (00133545, 00164966–00164969), 3♀ (00164970– 00164972) (USNM). Nevada: Nye Co.: Round Mountain, 38.717766 N 117.070256 W, 1920 m, 20 Apr 1915, E.G. Holt, 18 (00070116) (USNM). New Mexico: Guadalupe Co.: Milagro, 34.56186 N 105.14216 W, 1767 m, 23 Jun 1941, Burt Hogden, 28 (00117909, 00117927), 2♀ (00169993–00169994) (KU). Lincoln Co.: 5 mi N of Ruidoso, 33.38246 N 105.67296 W, 2225 m, 16 Aug 1970, J.R. and M.H. Sweet, 18 (00118223), 1♀ (00119073) (TAMU). San Miguel Co.: Las Vegas, Hot Springs, 35.653896 N 105.291116 W, H.S. Barber, 18 (00134360) (USNM). Sandoval Co.: Bandelier National Monument, 35.47186 N 106.1816 W, 2071 m, 23 Jun 1964, H.R. Burke, 18 (00134362) (USNM); 02 Jun 1969, W.F. Chamberlain, 38 (00119070–00119072) (TAMU). Jemez Springs, 35.768616 N 106.691676 W, 15 Jun 1915, Woodgate, 1♀ (00119069) (TAMU). Jemez Springs, 35.768616 N 106.691676 W, 1951 m, 07 Jun 1916 – 17 Jun 1916, unknown, 1♀ (00112154), paratype of junior synonym ( S. woodgatei ), 18 (00112153) (CNC), 18 (00119068) (TAMU), paratype of junior synonym ( S. woodgatei ), 18 (00122269) (USNM). Santa Fe Co. : Santa Fe , 35.686946 N 105.937226 W, 21 Jul 1926, E.C. Van Dyke, 1♀ (00242431) (CAS). Socorro Co. : Socorro County , 33.931946 N 106.948896 W, 18 Aug 1927, P.A. Readio, 18 (00112155) (CNC). Taos Co.: Taos, 36.407226 N 105.57256 W, 20 Aug 1927, P.A. Readio, 1♀ (00133546) (USNM). Torrance Co.: Moriarty, 34.59246 N 106.02576 W, 1896 m, 23 Jun 1941, L.H. Banker, 28 (00117922, 00117928), 3♀ (00109068–00109070) (KU). Tajique, 34.751676 N 106.287226 W, 25 Jun 1941, E.L. Todd, 38 (00117919–00117921), 1♀ (00109064) (KU); 25 Jun 1941, R.H. Beamer, 18 (00117794), 28 (00117917–00117918), 3♀ (00109065–00109067) (KU); 25 Jun 1941, Burt Hogden, 58 (00117912–00117916), 2♀ (00109062–00109063) (KU). North Dakota: Billings Co.: 8 mi W of Belfield at Fryburg Oil Field, 46.84996 N 103.40006 W, 776 m, 28 Jun 2000, T.J. Henry, Ericameria nauseosa , 38 (00122179, 00164962– 00164963), 2♀ (00134116, 00164964) (USNM). Stark Co.: 127 R Ave. 4 mi N of 48 R St. SW [9 mi S of South Heart], 46.733336 N 103.016666 W, 27 Jun 2000, T.J. Henry, Ericameria nauseosa , 18 (00164965) (USNM). South Dakota: Custer Co.: Custer, 43.766676 N 103.598336 W, 19 Aug 1927, H.H. Knight, paratype of junior synonym ( S. custeri ), 18 (00112152) (CNC), 1♀ (00134379) (USNM). Texas: Brewster Co.: Big Bend National Park, 29.565276 N 103.260556 W, 31 Aug 1986 – 02 Sep 1986, East, Haack, Kovarik, 1♀ (00119077) (TAMU). Big Bend National Park, Lost Mine Trail (lower), 29.276 N 103.278616 W, 1829 m, 09 Sep 1988, R. Anderson, 1♀ (00119078) (TAMU). Big Bend National Park, Pinnacles Trail (upper), 29.251666 N 103.296666 W, 2103 m, 21 Jun 2004, E.G. Riley, 1♀ (00245405) (TAMU). Jeff Davis Co.: H.O. Canyon, Davis Mountains, 30.68326 N 104.24326 W, 1890 m, 23 Aug 1968, J.E. Hafernik, 1♀ (00133547) (USNM). Madera Canyon , 30.55446 N 103.48496 W, 1031 m, 06 Aug 2000, E.G. Riley, 3♀ (00119074–00119076) (TAMU). Utah: Emery Co.: 6.2 mi W of Rt 24 in Temple Wash (on Goblin Valley Rd), 38.640446 N 110.662536 W, 1602 m, 19 Jun 1983, R.T. Schuh and M.D. Schwartz, Gutierrezia sarothrae , 28 (00107908, 00108398), 5♀ (00108399–00108403) (AMNH). Garfield Co.: Capitol Reef National Park, Grand Wash, Cobab Canyon Trail, 38.156 N 111.166676 W, 1827 m, 21 Jun 1983, R.T. Schuh, M.D. Schwartz, Chrysothamnus viscidiflorus , 18 (00107911), 3♀ (00108404– 00108406) (AMNH). San Juan Co.: 3 mi W of Clay Hills Crossing Road on Rt 263, T39S R15E, 37.295336 N 110.458786 W, 1524 m, 18 Jun 1983, R.T. Schuh and M.D. Schwartz, 1♀ (00108407), 18 (00107910) (AMNH). Goosenecks Overlook, Goosenecks State Park, 37.177786 N 109.925566 W, 1524 m, 17 Jun 1983, R.T. Schuh and M.D. Schwartz, Chrysothamnus sp. , 38 (00107904, 00108408, 00108410) (AMNH). Washington Co.: 15 mi N of Saint George, 37.326 N 113.586 W, 13 Jun 1935, Dobzhansky, 18 (00133544) (USNM). Wyoming: Crook Co.: Sundance, 44.406396 N 104.375286 W, 20 Jul 1927, H.H. Knight, paratype of junior synonym ( S. custeri ), 1♀ (00122244) (USNM). Sheridan Co.: Sheridan, 44.797226 N 106.955566 W, 03 Aug 1927, H.H. Knight, paratype, 18 (00134361) (USNM).

Figures

Plate 3. Dorsal habitus photographs of Slaterocoris punctatus—S. tibialis (see appendix 1 for USI codes and localities).

Map 9. Localities for Slaterocoris flavipes and S. sheridani.

Fig. 13. Hind leg coloration for Slaterocoris species.

Fig. 45. Slaterocoris species, endosoma. A–C. S. sheridani. A. 8 mi W of Belfield, ND. B. Englewood, CO. C. 12 mi W of Pueblo, CO. D, E. S. sparsus. D. Caliente Creek Road, CA. E. Sespe Canyon, CA.

Fig. 46. Slaterocoris sheridani, right paramere. A. Near Fish Creek, AZ. B. Sunflower, AZ. C, D. 1 mi E of Stoneman Lake, AZ. E. Temple Wash, UT. F. 8 mi W of Belfield, ND. G. 5 mi S of Clarkdale, AZ. H. Pueblo, CO. I. Englewood, CO. J. 36 mi SE of Globe, AZ. K. Mazatzal Wilderness Area, AZ. L. Tajique, NM.

Fig. 47. Slaterocoris sheridani, female genitalia, 8 mi W of Belfield, ND.

Fig. 2. One of three most parsimonious cladograms found in 40-taxa analysis for Slaterocoris and outgroups.