Sophiodela okinawana ( Nakane, 1957 ) Fukuda & Ogawa & Hori, 2019

Fukuda, Yuki, Ogawa, Ryo & Hori, Michio, 2019, The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia, Zootaxa 4661 (2), pp. 271-308 : 284

Sophiodela okinawana ( Nakane, 1957) , comb. nov.

( Figs 7C , 12C )

Cicindela (Sophiodela) chinensis okinawana Nakane, 1957:41 .

Cicindela (Sophiodela) okinawana: Fukuda et al. 2015:273 .

Specimens examined. (N = 1) [ Japan] 1♂ ( YFJ), Yona-Field, Yonakunigami-village , Okinawa, 17. III. 2015, Y. Fukuda leg.

Description. Internal sac everted dorsally in lateral view. DA largest in the bladders: DLL the second largest: VA as large as B: A exist: VLR, BLR and VB not exist.

VA short and roundly or triangular everted apically. DA everted basally. DA everted basally. B oval swollen right laterally, with well sclerotized spines, which are forming an oblong band. DLL swollen dorsally, covered numerously with well sclerotized spines, which are a forming band. Flagellum well short, curved and formed S shape. Medial tooth short, curved right ventro-laterally, tapered gently at apex.

Remarks. This species was treated as Cicindela (Sophiodela) okinawana by Fukuda et al. (2015), but the formal taxonomic treatment had not been accepted. As mentioned earlier, this species is clearly distinguished from the congener species of Sophiodela by the structures of the internal sac. Therefore, we reassigned S. okinawana in Sophiodela as an independent species.

Figures

FIGURE 7. Habitus of Sophiodela and Calochroa. A, Sophiodela chinensis chinensis (DeGeer) stat. nov.; B, Sophiodela japonica (Thunberg) stat. nov.; C, Sophiodela okinawana (Nakane) stat. nov.; D, Sophiodela ferriei (Fleutiaux) stat. nov.; E, Sophiodela chinensis flammifera (Horn) stat. nov.; F, Calochroa octonotata (Wiedemann); G, Calochroa carissima (Fleutiaux).

FIGURE 12. Everted internal sacs of Sophiodela. A, Sophiodela ferriei (Fleutiaux) stat. nov.; B, S. japonica (Thunberg) stat. nov.; C, S. okinawana (Nakane) stat. nov.; D, S. chinensis chinensis (DeGeer) stat. nov.; E, S. chinensis flammifera (Horn) stat. nov. Scales: 1.0 mm.