Stamnodes patamon ( Druce, 1893 )

Matson, Tanner A., 2023, A review of Mexican Stamnodes (Lepidoptera: Geometridae) with the description of 16 new species, European Journal of Taxonomy 911, pp. 1-79 : 49-51

publication ID	https://doi.org/ 10.5852/ejt.2023.911.2371
publication LSID	lsid:zoobank.org:pub:DB29E6F1-7925-46DB-8C9E-055C639203CE
DOI	https://doi.org/10.5281/zenodo.10376714
persistent identifier	https://treatment.plazi.org/id/03CEA053-3477-7873-FDFE-B0F2A869FCE5
treatment provided by	Plazi
scientific name	Stamnodes patamon ( Druce, 1893 )
status

Treatment

Stamnodes patamon ( Druce, 1893) View in CoL

Figs 24 View Figs 19–30 , 55 View Figs 55–57 , 94–95 View Fig View Fig

Marmopteryx patamon Druce, 1893: 178 View in CoL , ibidem 3: pl. 58 figs 7–8. Type locality: 6000 ft, Amula, Guerrero, Mexico. [NHMUK].

Stamnodes patamon View in CoL – Scoble 1999: 902 (catalogue). — Scoble & Hausmann 2007 (online catalogue). — Rajaei et al. 2022 (online catalogue).

Redescription

Druce’s original description of S. patamon did not include genitalic morphology. To his description, the following genitalic redescription and associated images are supplemented:

MALE GENITALIA ( Fig. 55 View Figs 55–57 ). Uncus long, narrow, slightly enlarged at apex. Inner face of valva with about five, long, sickle-shaped sclerotized ribbons originating from basal tubercle. Juxta cat-faced with stiff, crossing, submedial setae on posterior surface. Vesica with large spinose cornutus and adjacent patch of several hundred filamentous cornuti.

FEMALE GENITALIA. No material available for study.

Distribution

Mexico: Stamnodes patamon is primarily known to inhabit the pine-oak forests of the Northern Oaxaca Highlands and Sierra Madre del Sur, ranging north and westward into the Sierra Madre Occidental to at least western Durango. Reviewed material was often collected at elevations around 2000 m.

Biology

Stamnodes patamon is bi- or multivoltine, with adult records in March, April, May, and October. The immature stages of this taxon remain unknown, but its phylogenetic kin are mountain mahogany ( Cercocarpus ) feeders (Matson & Wagner in prep.).

Molecular characterization

Stamnodes patamon is represented in BOLD as BIN: BOLD:ABU6207 (n = 3). At present, the average pairwise intraspecific distance is 0.71%, the pairwise maximum intraspecific distance is 1.01%, and the distance to the nearest neighbour, Stamnodes sp. (?undescribed) (n = 1, Nuevo León), is 3.53%.

Remarks

During my review of this species, I came across individuals from the Sierra Madre Occidental that were located as far north and west as western Durango, which was surprising as I had previously considered S. patamon to be exclusive to the Sierra Madre del Sur. However, after examining the male genitalia of these individuals from Durango, I regard them as the same species.

A population that resembles S. patamon (S. nr patamon in Fig. 95 View Fig ) also extends into the northernmost areas of the Sierra Madre Oriental. Vitor Becker collected individuals in Gómez Farías, Tamaulipas, in May of 1997, and in Cerro Potosí, Nuevo León, in June of 1997. Unfortunately, I could only examine photographs of this material sent to me by Becker, as all but one specimen are held in his collection in Brazil, where regulations often prohibit biological material from leaving the country. However, one individual ( Fig. 23 View Figs 19–30 ) from Becker’s collection in Cerro Potosí is deposited at the USNM and has been barcoded. This specimen appears somewhat aberrant compared to most of the rest of Becker’s series and may represent yet another questionable entity. The COI barcode (BOLD Process ID: LNAUY180-19) was found to be 3.5% (pairwise) different from Oaxacan S. patamon . Although S. patamon from Durango, Guerrero, Oaxaca, and the putative S. nr patamon individuals from Nuevo León and Tamaulipas bear a subapical white patch on the forewing, this feature is lacking in the sequenced individual. The sequenced individual also has an overall lighter cream ground colour, and the medial transverse white patch on the hindwing underside appears straighter than the more arcing norm for S. patamon . Unfortunately, the sequenced individual is female, so I could not compare its genitalia to the entirely male series of S. patamon available for this study, nor to the putative individuals of S. nr patamon from Nuevo León and Tamaulipas in Becker’s collection in Brazil. For now, the sequenced individual will be regarded as the sole representative of a potentially undescribed species that warrants further investigation. In the Discussion and Figs 94 View Fig and 95 View Fig , it is listed as Stamnodes sp. (?undescribed).