Steginoporella cf. reingruberhohensis Zágoršek, 2003

Zágoršek, Kamil & Gordon, Dennis P., 2014, Revision of the Oligocene bryozoan taxa described by Stoliczka (1862), with the description of a new genus of Bryocryptellidae, Geodiversitas 36 (4), pp. 541-564 : 548-550

Steginoporella cf. reingruberhohensis Zágoršek, 2003

( Fig. 3A )

Ξ Steginoporella reingruberhohensis Zágoršek, 2003: 142 , pl. 15, figs 3, 4.

Non Cellaria haidingeri Reuss, 1848: 60 , pl. 7, fig. 30.

Eschara reussi Stoliczka, 1862: 88 .

Non Steginoporella cucullata Pouyet & David, 1979: 774 , fig. 3, pl. 3, fig. 10 (cum syn.).

MATERIAL EXAMINED. — Two syntypes of Eschara reussi registered as 1859/0026/0136. Ŋe chosen lectotype (designated here) is illustrated in Fig. 3A.

DIAGNOSIS. — Colony erect, columnar, multiserial, large with oval to circular transverse section. Autozooecia oval

to hexagonal, arranged in nine regular longitudinal rows. Mural rim wide and smooth.Cryptocyst shallow, perforated by 7-12 large pores but lacking opesiules. Orifice sunken, subcircular in shape, situated on the end of short peristome, bordered by a thin low rim. Ŋe vicarious avicularia (Bzooecia) up to twice as long and wide as autozooecia, with broad, truncate distal margin; cryptocyst perforated by about tenpores; palate wider than long, smooth, shallowly concave.Owing to poor preservation, opesial characters of vicariousaviculariaaresomewhatequivocalbutpolypidetube is well developed, its orifice similar to that of autozooecia.

REMARKS

Zágoršek (2003) synonymised this species with Steginoporella haidingeri (sensu Reuss 1848). SEM study of Stoliczka’s material, however, shows features more reminiscent of Steginoporella reingruberhohensis Zágoršek, 2003 . Ŋe main difference is the number of cryptocystal perforations. Whereas S. haidingeri usually has about 20-30 pores, S. reingruberhohensis has about 10-15, which is closer to the number (7- 12) in Eschara reussi . Owing to the unclear cryptocystal features of the vicarious avicularia, these two species cannot presently be synonymised and the status of Eschara reussi remains unclear.

David & Pouyet (1974) and Pouyet & David (1979) synonymised Eschara reussi Stoliczka, 1862 with Steginoporella cucullata (Reuss, 1848) . As already discussed by Zágoršek (2003), S. cucullata has a very wide mural rim, a consistently preserved pair of circular opesiules, and a wide vicarious avicularian palate. It is, moreover, a Miocene species.

Figures

FIG. 3. — A, Lectotype of Eschara reussi identified as Steginoporella cf. reingruberhohensis Zágoršek, 2003 showing large avicularia (top and lower right) and elevated distal zooecial rims; B-D, Adeonella ornatissima (Stoliczka,1862); B, general view of lectotype with autozooecia and maternal zooecia; C, maternal zooecia showing the characteristic semilunar aperture and autozooecia with a deep narrow sinus; D, lectotype designated here as older stage of colony development showing secondary calcification producing a transverse apertural bar; E, F, Adeonella mortisaga (Stoliczka, 1862), n. comb.; E, general view of lectotype showing autozooecia with two suboral avicularia and large maternal zooecia; F, maternal zooecium with very large aperture (partly broken) and autozooecia with a narrow sinus. Scale bars: A, B, D, E, 1 mm; C, F, 100 μm.