Heresiarches WESMAEL

Riedel, M., 2013, Contribution to the Ichneumoninae (Hymenoptera, Ichneumonidae) of Southeastern Asia: 2. Tribe Heresiarchini, Linzer biologische Beiträge 45 (2), pp. 2025-2076 : 2061-2063

Key of the known species of Heresiarches WESMAEL

(adopted from HEINRICH 1970: 73-75, modified, without Stenodontus spilocephalus CAMERON from India)

1 Mesosoma partly or more or less reddish............................................................................2

- Mesosoma black..................................................................................................................4

2 Mesosoma including the mesoscutum reddish. Metasoma red, tergites 1-3 with yellowish posterolateral marks.Area superomedia longer than wide, body length 13 mm. Male unknown. Java ..................................................... H. lieftincki HEINRICH 1934

- Mesosoma partly reddish, mesoscutum blackish, with yellowish stripes............................3

3 Mesoscutum blackish, with two yellow longitudinal stripes. Mesosoma ventrally and propodeum red, upper part of pleuras and area posteromedia black. Flagellum tricoloured. Body length 12 mm. India................................... H. rufus (CAMERON 1902)

- Mesoscutum blackish, with two median and two lateral yellowish stripes. Mesosoma, metasoma and coxa tricoloured: Black, reddish and yellow. Flagellum black. Probably the of H. rufus (CAMERON) . Laos......................................... H. rufomaculatus nov.sp.

4 Metasoma partly or completely red.....................................................................................5

- Metasoma black, with more or less extended pale bands or marks...................................13

5 Mesoscutum with two median longitudinal stripes and prescutellar carina yellow or ivory. First flagellomere slender in, about 6x as long as wide........................................6

- Mesoscutum without yellow longitudinal stripes................................................................7

6 Femur III yellowish-red, with black apex. Mesoscutum with ivory median stripes. Metasoma except first tergitered, slender, second tergite longer than wide, its lateral parts like the third tergite with fine and superficial puncture. In flagellum with 37 flagellomeres, widest flagellomere about 2x as wide as long. Tibia III infuscate apically. Body length 11 mm. India, Myanmar....................... H. himalayanus GUPTA 1955

- Femur III black. Mesoscutum with two yellowish median stripes and usually two lateral spots beneath tegula. Metasoma dark reddish-brown, tergits 1-5 darkened basally. Tergite 2 somewhat wider than long, like the third one with strong puncture. Flagellum with 37 flagellomeres, widest flagellomeres not wider than long. Body length 10 mm. Male unknown. Myanmar......................... H. annaelisae HEINRICH 1970

7 Propodeum completely black, sides of scutellum with narrow ivory stripes. Legs including coxa III and femur III yellowish-red, tibia III with infuscate apex. Metasoma mainly red, tergites 1-2 with yellow posterolateral marks, apical marks of tergites 6-7 yellow, central tergits darkened basally. Flagellum with 34 flagellomers. Body length 10 mm. Myanmar.................................................................. H. malaisei HEINRICH 1970

- Propodeum with large dorsolateral ivory spots. Scutellum with wide lateral ivory stripes or almost completely ivory......................................................................................8

8 Coxa III mainly red. Body length 15 mm. Scutellum completely yellow or with a black central longitudinal stripe. In flagellum with 39 flagellomeres, widest flagellomeres wider than long. Metasoma almost completely red, Yellowish are indistinct posterolateral spots on tergites 1-2 and apical spots on tergites 6-7. Tarsus III reddish. In metasoma black apically, flagellum black, tarsus III dark brown. Sulawesi ................ .......................................................................................... H. bonthainensis HEINRICH 1934

- Coxa III mainly blackish. Smaller species..........................................................................9

9 Metasoma with the three or four apical tergites completely or partly black.....................10

- Tergites 2-7 mainly red or reddish-brown.........................................................................12

10 Tibia III completely pale yellow. In tergites 1-3 yellowish-red with yellowish apical margins. Tergite 4 black, with frontal and apical margins partly reddish. In the (questionable) only postpetiolus and apical half of tergite 2 reddish. Body length 11 mm. Java ............................................................................. H. daedalus (TOSQUINET 1903)

- Tibia III infuscate apically................................................................................................11

11 In flagellum with 39 flagellomeres, without an ivory ring. Tergites 2-4 red, without apical yellowish spots, tarsus III infuscate. with 40 flagellomeres, the first one almost 6x as long as wide, widest flagellomeres not wider than long. Tarsus III yellowish-red. Petiolus and tergites 4-7 black. Body length 12-13 mm. India, Myanmar.... ................................................................................... H. pseudobirmanicus HEINRICH 1970

- flagellum with 33 flagellomeres, without ivory ring. Mesosternum and metapleurum without pale coloration. yellowish spots on tergites 1-4. Tarsus III blackish. unknown. Body length 10 mm. Vietnam.............. H. flavomaculatus nov.sp.

12 Larger species, body length 15 mm. Flagellum blackish, with yellowish stripe. Coxa III completely black. Petiolus blackish, postpetiolus and all tergites reddish, without pale markings. Femur III red, Tarsus III yellowish. Taiwan... H. heinrichi UCHIDA 1932

- Smaller species, body length 10 mm. Flagellum completely black. Coxa III ivory dorsally. Mesoscutum and Propodeum very short, metasoma wide. Tergites 2-7 pale reddish. Femur III and tibia III reddish, sometimes infuscate apically. Tarsus III reddish or blackish. Flagellum with 35 flagellomeres. Myanmar......................................... ............................................................................................. H. rotundus HEINRICH 1970

13 Metasoma black, only tergites 6-7 with ivory coloration. Male unknown. Korea................ ................................................................................................ H. coreanus UCHIDA 1926

- Metasoma black, with more or less extended pale bands or marks...................................14

14 Coxal carina lacking, tergites 2-5 with bluntly extended posterolateral corners and with posterolateral ivory spots. Flagellum with 35 flagellomeres, first one about 4x as long as wide, widest flagellomeres as wide as long. Mesoscutum black, prescutellar carina yellowish. Body length 10 mm. unknown. Nepal............... H. atanyjoppoides nov.sp.

- Coxal carina present, tergites with rounded posterolateral corners...................................15

15 All tergites with yellow or whitish spots or bands............................................................16

- At least tergite 5 completely blackish, without pale coloration.........................................21

16 Coxa III mainly or partly reddish, at least ventrally. Tarsus III reddish, brownish or black..................................................................................................................................17

- Coxa III mainly black. Femur III red, apically infuscate, or completely black. Tarsus III black.............................................................................................................................19

17 Coxa III bicoloured: red, with yellowish spot dorsal. Femur III red, tarsus III brownish. In flagellum strongly lanceolate, with 34 flagellomeres, widest flagellomeres almost 2 times as wide as long. Ivory stripe on flagellomeres 4-15. In flagellum black. Body length 10 mm. Sulawesi .................................................. H. proximus HEINRICH 1934

- Coxa III tricoloured: red, with yellowish and black spots dorsally. In flagellum less strongly lanceolate, widest flagellomeres 1.5x as wide as long........................................18

18 Femur III red, tarsus III reddish-brown. Flagellum very slightly lanceolate, with 38 flagellomeres, first flagellomere 3.5x as long as wide, widest flagellomeres 1.2x as wide as long. Ivory stripe on flagellomeres 7-13. Body length 10.5 mm. unknown. India..................................................................................................... H. indicus nov.sp.

- Femur III red, infuscate apically. Tarsus III black. In flagellum slightly lanceolate, with 32 flagellomere, first flagellomere 3.2x as long as wide, widest ones 1.5x as wide as long. In flagellum with 30 flagellomeres. Body length 9-9.5 mm. Taiwan.................. .......................................................................................................... H. formosanus nov.sp.

19 Flagellum of with ivory stripe or ring. Tyloids wide-oval, on flagellomeres 9-18. Body length 9 mm. unknown. Vietnam........................................... H. tyloidus nov.sp.

- Flagellum of black. Tyloids long-oval or linear............................................................20

20 Femur III red, black apically. Tarsus III blackish. Body length 8-10 mm. In first flagellomere 6x as long as wide, flagellum with 32-35 flagellomers. Myanmar................... ................................................................................................. H. secundus HEINRICH 1970

- Femur III and tarsus III completely black. Body length 10 mm ............................................ ............................................................ H. megaleudoxius HEINRICH 1970 (questionable)

21 Coxa III mainly red. Tarsus III cream-yellow...................................................................22

- Coxa III mainly black........................................................................................................23

22 Coxa III reddish. Tergites 3-5 without pale coloration. Flagellum with 36 flagellomeres, first one about 6x as long as wide, widest flagellomeres not wider than long. Ovipositor sheath very long, as long as tergites 6 and 7 together. Body length 10 mm. unknown. Myanmar............................................... H. cingulatus HEINRICH 1970

- Coxa III red, with dorsal ivory spot. In flagellum with 44 flagellomeres, first one 3.4x as long as wide, widest flagellomeres about1.6x as wide as long. Area superomedia with complete carination. Ovipositor sheath as long as tergite 7. Body length 9 mm. Vietnam....................................................................... H. rufocoxatus nov.sp.

23 Femur III black..................................................................................................................24

- Femur III red, at least in the basal half..............................................................................25

24 Smaller, body length 8-10 mm. Tarsus III infuscate. Tergits 1-3 with small ivory dorsolateral marks, tergites 6 and 7 with ivory apical bands. Europe ................................... .............................................................................................. H. eudoxius (WESMAEL 1845)

- Larger, body length 14-15 mm. Tarsus III cream-yellow. Postpetiolus with yellow dorsolateral spot, tergites 2-3 with wide yellowish or reddish-yellow apical bands, following tergites with narrow yellowish apical margins. unknown. Laos, Vietnam........ ........................................................................................................... H. tarsator nov.sp.

25 Tarsus III yellow-red in and cream-yellow in. Flagellum with 32-34 flagellomeres. Tergites 1 and 2 with yellowish dorsolateral marks or apical bands. Tergite 6 usually with an apical ivory band, tergite 7 mainly ivory. Ovipositor sheath in short, shorter than tergite 7. Body length 9-10 mm. Java, Laos.................................... ......................................................................................................... H. felix HEINRICH 1934

- Tarsus III blackish. Metasoma with more extensive yellowish or ivory coloration..........26

26 Larger species, body length 13 mm. Flagellum with about 40 flagellomeres, first one about 6x as long as wide, widest flagellomeres slightly wider than long. Malar space ivory. Tergites 1-4 with triangular yellowish dorsolateral marks, tergite 6 with yellowish apical margin, tergite 7 with a small yellowish spot. Myanmar........................... .................................................................................... H. megaleudoxius HEINRICH 1970

- Smaller species, body length 6-7 mm. Coloration ivory, not yellowish. Tergite 7 mainly ivory. Malar space with blackish stripe.................................................................27

27 In flagellum with 29 flagellomeres, very slender, the first one about 7x as long as wide, widest flagellomere slightly wider than long. Tergite 1-2 with wide apical ivory bands, apical band of third tergite interupted in the middle, tergite 5 with narrow apical band, tergite 6-7 mainly ivory. unknown. Myanmar......................................................... .......................................................................................... H. albicinctus HEINRICH 1970

- flagellum with 27 flagellomeres. Tergites 1-2 with dorsolateral ivory spots, apical bands of tergite 6 and tergite 7 mainly ivory. Tergites 3-5 black. unknown. Taiwan....... ................................................................................................................... H. minor nov.sp.