Sticholotis quadriguttata (Gorham)

Tomaszewska, Wioletta & Łączyński, Piotr, 2010, A contribution to Sticholotis Crotch from Myanmar (Coleoptera: Coccinellidae: Sticholotidini), Zootaxa 2658 (1), pp. 51-68 : 65

Sticholotis quadriguttata (Gorham)

( Figs. 2 , 56–67 )

Orcus quadriguttatus Gorham, 1895: 689 .

Sticholotis quadriguttata: Łączyṅski & Tomaszewska, 2009: 609 .

Diagnosis. This species is easily distinguished among species of Sticholotis from Myanmar by its small body and two large, brown maculae on each elytron.

Redescription. Female. Length 2.1 mm; TL/EW = 1.15; PL/PW = 0.47; EL/EW = 0.85; GD/TL = 0.5.

Body ( Figs. 2 , 56 ) rounded, strongly convex; pronotal margins very narrow, visible from above; elytral margins comparatively explanate, entirely visible from above. Pronotum, scutellum and background of elytra brownish black; head and labrum dark reddish brown. Each elytron with two large, round, reddish brown spots along mid line of disk, narrowly connected to each other; elytral and pronotal lateral margins reddish brown. Punctures on pronotum 1.5–2.5 diameters apart, rather fine and moderately deep; punctures on elytra slightly coarser and deeper than those on pronotum, 1.0–2.0 diameters apart; surfaces between elytral and pronotal punctures polished and shiny; dorsum apparently glabrous. Ventral surface dark reddish brown with meso- and metaventrite, especially along middle and anterior part of abdominal ventrite I, infuscate. Ventral mouthparts and antennae yellowish brown; legs slightly darker.

Head ( Figs. 57, 59 ) flat medially, finely punctate and moderately densely setose. Clypeus weakly arcuate anteriorly, scarcely reflexed at anterior edge. Eyes moderately large, rather coarsely faceted, dorsally separated by about 3 times eye width; interocular distance nearly 0.65 times head width; inner margins of eyes slightly rounded, convergent anteriorly. Maxilla ( Figs. 58, 59 ) with terminal palpomere about 2.3 times longer than wide, parallel-sided along basal half of its length, strongly obliquely truncate towards apex; labial terminal palpomere narrowed and acuminate, distinctly narrower than penultimate palpomere. Antenna ( Fig. 58 ) 11-segmented with narrow 3-segmented club.

Prothorax ( Figs. 57, 62 ) about 0.9 times base of elytra; pronotum with groove extending parallel along base, curving anteriorly near hind angles; pronotal hypomeron and prosternum smooth; anterior lobe of prosternum distinctly bordered with anterior edge emarginate medially; prosternal process ( Figs. 59, 60 ) subtruncate at apex, without distinct lateral carinae, moderately coarsely and rather sparsely punctate, punctures with moderately long setae. Mesoventral intercoxal process ( Fig. 61 ) about 1.35 times mesocoxal diameter. Metaventrite with complete discrimen, moderately coarsely and sparsely punctate; postcoxal lines curved and complete. Elytral epipleuron broad with maximum width at metaventrite, narrowing posteriorly but complete to apex, without distinct foveae. Wings well-developed.

Abdomen ( Figs. 63 , 64 ) with 5 ventrites; ventrite I along mid line about 3.0 times longer than ventrite II; postcoxal line of first ventrite curved posteriorly and laterally, closely paralleling posterior margin, incomplete laterally; postcoxal disk microreticulate and sparsely punctate ( Figs. 63 , 65 ); ventrite V triangularly produced posteriorly; abdominal segment VIII in female as in Fig. 66 .

Female genitalia as in Fig. 67 . Proctiger (T10) well developed, somewhat triangular in shape; ovipositor sclerotized, coxites triangularly oval, entirely separated; styli absent; spermatheca not observed.

Male not known.

Material examined. Types. Lectotype (here designated), female: “ Rangoon Birmania, Fea V. 1885 / Typus / quadriguttatus Gorh. / Orcus quadriguttatus / O. quadriguttatus Gorh. typus!/ Syntypus, Orcus quadriguttatus Gorham, 1895 / Museo Civico di Genova 1895” ( MCSN).

Note. The designation of the lectotype for O. quadriguttatus Gorham, 1895 , is made to fix the taxonomic status of this species.

Figures

FIGURES 1–5. Habitus, dorsal view; 1) Sticholotis bipunctata (Gorham); 2) Sticholotis quadriguttata (Gorham); 3) Sticholotis discoidea (Gorham); 4) Sticholotis dohrni Weise; 5) Sticholotis carinica (Gorham).

FIGURES 56–63. Sticholotis quadriguttata (Gorham); 56) body, lateral view; 57) head, prothorax and base of elytra, antero-dorsal; 58) antenna, maxillary palp and prothoracic hypomeron; 59) head and prothorax, ventral; 60) prosternal process; 61) meso-, metathorax, and abdominal ventrite I, ventral; 62) pronotum, lateral; 63) abdomen, female, ventral.

FIGURES 64–67. Sticholotis quadriguttata (Gorham); 68–71. Sticholotis carinica (Gorham); 64, 68) abdomen, ventral; 65, 69) abdominal postcoxal line; 66, 70) abdominal segment VIII, female, ventral; 67, 71) female genitalia.