Tectarioideae, sensu Christenhusz & Chase, 2014

Wang, Fa-Guo, Barratt, Sam, Falcón, Wilfredo, Fay, Michael F., Lehtonen, Samuli, Tuomisto, Hanna, Xing, Fu-Wu & Christenhusz, Maarten J. M., 2014, On the monophyly of subfamily Tectarioideae (Polypodiaceae) and the phylogenetic placement of some associated fern genera, Phytotaxa 164 (1), pp. 1-16 : 8

Monophyly of Tectarioideae

Our results are in line with the suggestion that Tectarioideae sensu Holttum (1947) are polyphyletic. The placement of Cyclopeltis in subfamily Lomariopsidoideae ( Smith et al. 2006, Schuettpelz & Pryer 2007) is supported by our analyses, as is the inclusion of Dryopsis and Ctenitis in Dryopteridoideae ( Li & Lu 2006, Liu et al. 2007). Indeed, Dryopsis has already been merged with Dryopteris ( Zhang 2012) , although this may cause Dryopteris not to be monophyletic ( Christenhusz & Chase 2014). Ctenitis (including Ataxipteris ) forms a lineage with Pseudotectaria and both are deeply embedded in Dryopteridoideae , even though these three genera share morphological characteristics with Tectarioideae and were associated with it previously. Pleocnemia , which was placed in Tectariaceae by Smith et al. (2006) and Christenhusz et al. (2011), is here shown to belong to Dryopteridoideae , agreeing with other recent studies (e.g. Liu et al. 2014). With the exclusion of these genera, Tectarioideae are monophyletic and are composed of Arthropteris , Pteridrys , Triplophyllum , Hypoderris and Tectaria . We were not able to sample Aenigmopteris , so its placement remains uncertain.