Tetrapleurus heteroderes, Fauvel, 1902

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1067-1806 : 1067-1806

heteroderes Fauvel, 1902 , see: Tetrapleurus .

kauaiensis Sharp, 1908: 541 ( Lispinodes ; Type locality: Kauai, Koholuamano).

— Bernhauer and Schubert, 1910: 20 ( Lispinodes ; catalog).

— Nishida, 1994: 72 ( Lispinodes ; Hawaii).

— Nishida, 1997: 61 ( Lispinodes ; Hawaii).

- DISTRIBUTION: Hawaii.

mauiensis Sharp, 1908: 541 ( Lispinodes ; Type locality: Maui, Haleakala, 5000 ft.).

— Bernhauer and Schubert, 1910: 20 ( Lispinodes ; catalog).

— Nishida, 1994: 72 ( Lispinodes ; Hawaii).

— Nishida, 1997: 61 ( Lispinodes ; Hawaii).

- DISTRIBUTION: Hawaii.

molokaiensis Sharp, 1908: 540 ( Lispinodes ; Type locality: Molokai).

— Bernhauer and Schubert, 1910: 20 ( Lispinodes ; catalog).

— Nishida, 1994: 72 ( Lispinodes ; Hawaii).

— Nishida, 1997: 61 ( Lispinodes ; Hawaii).

- DISTRIBUTION: Hawaii.

obscurus Sharp, 1908: 541 ( Lispinodes ; Type locality: Molokai, mts. 4500 ft.).

— Bernhauer and Schubert, 1910: 20 ( Lispinodes ; catalog).

— Nishida, 1994: 72 ( Lispinodes ; Hawaii).

— Nishida, 1997: 61 ( Lispinodes ; Hawaii).

- DISTRIBUTION: Hawaii.

oxytelinus Sharp, 1908: 540 ( Lispinodes ; Type locality: Lanai).

— Bernhauer and Schubert, 1910: 20 ( Lispinodes ; catalog).

— Nishida, 1994: 72 ( Lispinodes ; Hawaii).

— Nishida, 1997: 61 ( Lispinodes ; Hawaii).

- DISTRIBUTION: Hawaii.

pallescens Blackburn, 1885 , see: Clavilispinus .

pallidus Sharp, 1908: 540 ( Lispinodes ; Type locality: Lanai, 2000 ft.).

— Bernhauer and Schubert, 1910: 20 ( Lispinodes ; catalog).

— Nishida, 1994: 72 ( Lispinodes ; Hawaii).

— Nishida, 1997: 61 ( Lispinodes ; Hawaii).

- DISTRIBUTION: Hawaii.

quadratus Blackburn, 1885 , see: Lispinus .

rhodesianus Bernhauer, 1934 h: 484 ( Lispinodes ; Type locality: N. W. Rhodesia: Shigariatombwes. Also cited from Belgian Congo).

— Scheerpeltz, 1956a: 514 ( Lispinodes ; characters; Uganda; Kenya; Tanganyika; Belgian Congo). - DISTRIBUTION: Uganda, Kenya, Tanzania, Congo, Zambia.

rugosus Fauvel, 1902 , see: Tetrapleurus .

seriatus Fauvel, 1902 , see: Tetrapleurus .

tenuipunctus Bernhauer, 1932a: 73 ( Lispinodes ; Type locality: Bas-Uele: Koteli).

— Scheerpeltz, 1933: 1010 ( Lispinodes ; catalog).

— Scheerpeltz, 1956a: 515 ( Lispinodes ; characters; French Congo; Belgian Congo).

- DISTRIBUTION: French Equatorial Africa, Congo.

Parespeson

[3 species; Oriental Region]

Parespeson Bernhauer, 1926d: 261 (subgenus of Espeson ; species included: angustissimus ). TYPE SPECIES: Espeson (Parespeson) angustissimus Bernhauer , fixed by monotypy. — Blackwelder, 1943: 144 (subgenus of Espeson ; type species angustissimus ).

— Blackwelder, 1952: 294 (type species: angustissimus ).

— Scheerpeltz, 1970: 126, 127 (valid genus; key to species).

angustiformis Cameron, 1950d: 3 ( Espeson ; subgenus Parespeson ; Type locality: Selangor: The Gap).

— Scheerpeltz, 1970: 128 ( Parespeson ; characters; Malaysia).

- DISTRIBUTION: Malaysia.

angustissimus Bernhauer, 1926d: 261 ( Espeson ; subgenus Parespeson ; Type locality: Luzon, Provinz Ilocos Norte, Bangui).

— Scheerpeltz, 1933: 1020 ( Espeson ; subgenus Parespeson ; catalog).

— Scheerpeltz, 1970: 128 ( Parespeson ; characters; Philippines).

- DISTRIBUTION: Philippines.

lombokianus Cameron, 1945a: 141 ( Espeson ; subgenus Parespeson ; Type locality: Lombok: Sapit, altitude 2000 m).

— Scheerpeltz, 1970: 127 ( Parespeson ; characters; Lombok).

- DISTRIBUTION: Indonesia.

Pseudespeson

[12 species; Neotropical, Palaearctic, Ethiopian, Madagascan, Oriental] Pseudespeson Lecoq, 1994: 299 (species included: rossii , crassulus , crenicollis , gigantulus , indomalayicus, luzonicus , nepalicus , nitens , philippinus , philippinus curtus , and scotti ). TYPE SPECIES: Pseudespeson rossii Lecoq , fixed by original designation.

— Drugmand, 1998a: 223 (list of types in l’Institut royal des Sciences naturelles de Belgique). crassulus Fauvel, 1902: 35 ( Espeson ; Type locality: Antilles: Guadeloupe; Saint Vincent).

— Bernhauer and Schubert, 1910: 24 ( Espeson ; catalog).

— Blackwelder, 1943: 144 ( Espeson ; characters; Montserrat; Guadeloupe; St. Lucia; St. Vincent). — Scheerpeltz, 1970: 116 ( Espeson ; characters; Montserrat; Guadeloupe; Martinique; Santa Lucia; St. Vincent; Grenada).

— Lecoq, 1994: 304 ( Pseudespeson ).

- DISTRIBUTION: West Indies.

crenicollis Fauvel, 1878f: 196 ( Thoracophorus ; Type locality: Iles Key).

— Fauvel, 1879: 75 ( Thoracophorus ; Gilolo).

— Bernhauer and Schubert, 1910: 24 ( Espeson ; catalog).

— Scheerpeltz, 1970: 121 ( Espeson ; characters; Key Island; Halmahera Island [Gilolo]).

— Lecoq, 1994: 304 ( Pseudespeson ).

- DISTRIBUTION: Indonesia.

curtus Bernhauer, 1926 , see: philippinus Bernhauer, 1926 .

gigantulus Bernhauer, 1915d: 97 ( Espeson ; Type locality: Deutsch-Ostafrika: West-Usambara ).

— Scheerpeltz, 1933: 1019 ( Espeson ; catalog).

— Scheerpeltz, 1970: 125 ( Espeson ; characters; Tanzania).

— Lecoq, 1994: 303, 304 ( Pseudespeson ; characters).

- DISTRIBUTION: Tanzania.

gomyi Lecoq, 1995: 416 ( Pseudespeson ; Type locality: Moihani , Mohéli, archipel des Comores).

- DISTRIBUTION: Comoros.

indomalayensis Bernhauer, 1926d: 262 ( Espeson ; Type locality: Polillo).

— Scheerpeltz, 1933: 1019 ( Espeson ; catalog).

— Scheerpeltz, 1970: 122 ( Espeson ; characters; Philippines).

— Lecoq, 1994: 304 ( Pseudespeson ; cited as indomalayicus).

- DISTRIBUTION: Philippines.

luzonicus Bernhauer, 1926d: 262 ( Espeson ; Type locality: Luzon, Provinz Nueva Vizcaya, Imugan ).

— Scheerpeltz, 1933: 1019 ( Espeson ; catalog).

— Scheerpeltz, 1970: 123 ( Espeson ; characters; Philippines). — Lecoq, 1994: 304 ( Pseudespeson ).

- DISTRIBUTION: Philippines.

minusculus Coiffait, 1982 , see: nepalicus Coiffait, 1981 .

nepalicus Coiffait, 1981d: 334 ( Espeson ; Type locality: Royal Chitwan, National Park, Theral; [Note: The preceding locality is as cited on page 325 and is a correction of the citation on page 334, “Royal Shitwan, National par, Theral ”]).

— Lecoq, 1994: 304 ( Pseudespeson ).

- DISTRIBUTION: Nepal.

minusculus Coiffait, 1982: 135 ( Bothrys ; Type locality: Nepal: Siwalik , 660 m).

— Lecoq, 1994: 304 ( Pseudespeson ; synonym of nepalicus ).

nitens Fauvel, 1902: 36 ( Espeson ; Type locality: Bresil; Guadeloupe).

— Bernhauer and Schubert, 1910: 24 ( Espeson ; catalog).

— Blackwelder, 1943: 147 ( Espeson ; characters; Guadeloupe).

— Scheerpeltz, 1970: 117 ( Espeson ; characters; Guadeloupe; Martinique; Grenada; Trinidad; Venezuela; Brazil).

— Lecoq, 1994: 304 ( Pseudespeson ).

— Lecoq, 1994: 304 ( Pseudespeson ).

- DISTRIBUTION: Brazil, Venezuela, West Indies.

orousseti Lecoq, 1995: 417 ( Pseudespeson ; Type locality: Philippines, Luzon , Sagada).

- DISTRIBUTION: Philippines.

philippinus Bernhauer, 1926d: 262 ( Espeson ; Type locality: Luzon, Provinz Laguna,

Los Baños. Siargao, Dapa. Masbate, Aroroy. Dinagat. Mindanao, Provinz Lanao, Mumungan).

— Scheerpeltz, 1933: 1019 ( Espeson ; catalog).

— Scheerpeltz, 1970: 123 ( Espeson ; characters; Philippines).

— Lecoq, 1994: 304 ( Pseudespeson ).

- DISTRIBUTION: Philippines.

curtus Bernhauer, 1926d: 262 ( Espeson ; variety of philippinus ; Type locality: Dapa).

— Scheerpeltz, 1933: 1019 ( Espeson ; synonym of philippinus ).

— Scheerpeltz, 1970: 124 ( Espeson ; valid species; characters; Philippines).

— Lecoq, 1994: 304 ( Pseudespeson ; subspecies of philippinus ).

rossii Lecoq, 1994: 302 ( Pseudespeson ; Type locality: Sierra Leone: Western Area, Number Two River Forest . Also cited from Ivory Coast and Guinea).

- DISTRIBUTION: Guinea, Sierra Leone, Ivory Coast.

scotti Bernhauer, 1922b: 166 ( Espeson ; Type locality: Seychellen: Silhouette : Umgebung von Pot-à-eau, circa 1500 ft.).

— Scheerpeltz, 1933: 1020 ( Espeson ; catalog).

— Scheerpeltz, 1970: 125 ( Espeson ; characters; Seychelles).

— Lecoq, 1994: 304 ( Pseudespeson ).

- DISTRIBUTION: Seychelles.

Synaenictus

[1 species; Ethiopian Region]

Synaenictus Patrizi, 1947: 228 (species included: foveicauda ). TYPE SPECIES: Synaenictus foveicauda Patrizi , fixed by monotypy.

— Blackwelder, 1952: 367 (type species: foveicauda ).

— Seevers, 1965: 341 (characters; may be Pselaphidae not Staphylinidae ; myrmecophilus species).

— Burakowski and Newton, 1992: 38 (uncertain placement in Osoriinae ).

foveicauda Patrizi, 1947: 229 ( Synaenictus ; Type locality: Elmenteita , Kenya Colony).

- DISTRIBUTION: Kenya.

Subtribe Lispinina

Lispinina Bernhauer and Schubert, 1910 .

— Arnett, 1963: 262 (characters; list of North American genera).

— Newton and Thayer, 1992: 59 (subtribe of Thoracophorini ).

— Newton, Thayer, Ashe, and Chandler, 2000: 377 (characters; notes).

Lispini Bernhauer and Schubert, 1910: 19 (genera included: Ancaeus , Lispinodes , Holosus , Lispinus ). TYPE GENUS: Lispinus (see Newton and Thayer, 1992: 59).

— Scheerpeltz, 1933: 1009 (world catalog supplement).

— Blackwelder, 1942: 77, 78 (characters; key to genera and subgenera).

— Blackwelder, 1944: 107 (checklist of species from Mexico, Central and South America, and the West Indies).

Lispininae Bernhauer and Schubert, 1910.

— Moore, 1964a: 87 (characters; notes).

— Moore and Legner, 1974b: 77 (characters; key to genera of America north of Mexico).

— Moore and Legner, 1974g: 555 (key to genera of America north of Mexico).

— Moore and Legner, 1975: 166 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 143 (characters; tabular key to genera of North America north of Mexico).

— Downie and Arnett, 1996: 426 (characters; key to genera and species of Northeastern

North America).

Lispinini Bernhauer and Schubert, 1910.

— Blackwelder, 1942: 76, 78 (characters; key to subtribes).

— Blackwelder, 1944: 107 (checklist of species from Mexico, Central and South America, and the West Indies).

— Hatch, 1957: 243 (characters; key to species of Pacific Northwest).

— Arnett, 1963: 242, 262 (key to North American genera; characters; list of North American genera).

Lispinus

[282 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]

Lispinus Erichson, 1839b: 31 (without included species, for first included species see Erichson, 1840). TYPE SPECIES: Lispinus attenuatus Erichson , fixed by subsequent designation by Duponchel, 1841: 57.

— Erichson, 1840: 828 (characters; first included species: attenuatus , laticollis , striola , angustatus , linearis , tenellus , exiguus ).

— Lacordaire, 1854: 126 (characters; notes; list of species).

— LeConte, 1861a: 72 (characters).

— LeConte, 1863: 27 (list of North American species).

— Fauvel, 1864b: 40 [= 1865: 44] (characters; list of and key to species).

— Gemminger and Harold, 1868: 677 (catalog).

— Sharp, 1876: 411 (notes).

— LeConte, 1877: 250 (key to species).

— Fauvel, 1878d: 176 [= 1878e: 12] (key to North American species).

— Fauvel, 1878f: 33 (characters; key to New Guinea species).

— Duvivier, 1883: 204 (catalog).

— Sharp, 1887: 781 (notes; Mexican and Central American species).

— Fauvel, 1897d: 240 (catalog of species of Madeira).

— Bernhauer and Schubert, 1910: 21 (world catalog; 94 species).

— Blatchley, 1910: 482 (characters).

— Leng, 1920: 93 (catalog of North American species).

— Cameron, 1921a: 350, 399 (characters; key to Singapore species; catalog).

— Winkler, 1925: 323 (catalog for Palaearctic region).

— Cameron, 1925: 7, 103 (catalog of Indian species).

— Scheerpeltz, 1933: 1012 (world catalog supplement).

— Bernhauer, 1934h: 483 (key to species of South Africa).

— Paulian, 1941: 139 (larval characters).

— Blackwelder, 1942: 83 (characters; notes).

— Blackwelder, 1943: 129 (characters; type species: attenuatus ; key to species of West Indies).

— Blackwelder, 1944: 107 (checklist of species from Mexico, Central and South America, and the West Indies).

— Shibata, 1973c: 22 (checklist of species of Taiwan).

— Tikhomirova, 1973a: 142 (checklist of species of USSR).

— Moore and Legner, 1975: 168 (catalog; North America north of Mexico).

— Shibata, 1976: 82 (checklist of species of Japan).

— Moore and Legner, 1979: 146 (characters; notes).

— Hammond, 1984: 196 (checklist of species of Borneo).

— Newton, 1990: 92 (characters in key; three described Australian species).

— Irmler, 1994: 54 (key to Neotropical species).

— Naomi, 1996b: 762 (characters; species of Japan).

— Drugmand, 1998a: 212 (list of types in l’Institut royal des Sciences naturelles de Belgique).

— Newton, Thayer, Ashe, and Chandler, 2000: 377 (2 Nearctic species; notes; characters in key).

Paralispinus Eichelbaum, 1913: 117 (subgenus of Lispinus ; species included: amaniensis ). TYPE SPECIES: Lispinus (Paralispinus) amaniensis Eichelbaum , fixed by monotypy.

— Blackwelder, 1952: 290 (subgenus of Lispinus ; type species: amaniensis ).

Pseudolispinodes Bernhauer, 1926d: 258 (species included: longipennis , madurensis , latiusculus , sinuatus ). TYPE SPECIES: Holosus madurensis Bernhauer, 1914 , fixed by subsequent designation by Blackwelder, 1942: 88.

— Cameron, 1930: 65 (characters).

— Scheerpeltz, 1933: 1010 (world catalog supplement).

— Blackwelder, 1943: 120 (characters; [Note: Species included in Pseudolispinodes by Blackwelder are now in Nacaeus ]; type species: madurensis ; key to species of West Indies).

— Blackwelder, 1952: 327 (subgenus of Lispinus ; type species: madurensis ).

— Abdullah and Qadri, 1970: 114 (key to species of West Pakistan).

— Drugmand, 1998a: 223 (list of types in l’Institut royal des Sciences naturelles de Belgique).

Spinilus Blackwelder, 1942: 83 (subgenus of Lispinus ; species included: bistriatus , sinuatus ). TYPE SPECIES: Lispinodes bistriatus Fauvel , fixed by original designation.

— Blackwelder, 1952: 355 (synonym of Lispinus ; type species).

Aulacolispinus Scheerpeltz, 1957c: 223 , 239, 240 (subgenus of Lispinus ; species included: unistriatus , lineipennis , cheesmani , sutteri , sulcipennis , macropterus , sulcatus , validicornis , sulcatulus , germanus , striola ). TYPE SPECIES: Lispinus unistriatus Fauvel , fixed by original designation.

Stictolispinus Scheerpeltz, 1957c: 241 (subgenus of Lispinus ; species included: quadripunctulus ). TYPE SPECIES: Lispinus quadripunctulus Fauvel , fixed by original designation.

abdominalis Bernhauer, 1929 i: 355 ( Lispinus ; Type locality: Luzon, Haight’s Place).

— Scheerpeltz, 1933: 1012 ( Lispinus ; catalog).

- DISTRIBUTION: Philippines.

acicularis Fauvel, 1904b: 80 ( Lispinus ; Type locality: Java).

— Bernhauer and Schubert, 1910: 21 ( Lispinus ; catalog).

— Scheerpeltz, 1933: 1012 ( Lispinus ; catalog).

- DISTRIBUTION: Indonesia.

aciculatus Bernhauer, 1927: 47 ( Lispinus ; Type locality: Gazaland: Chirinda).

— Bernhauer, 1934h: 483 ( Lispinus ; characters in key).

- DISTRIBUTION: South Africa.

acupunctus Eppelsheim, 1895a: 137 ( Lispinus ; Type locality: Gabon).

— Fauvel, 1903e: 235 ( Lispinus ; Cameroon; Togo; Gabon; Usambara).

— Paulian, 1942: 347 ( Lispinus ; larval characters).

— Fagel, 1963a: 339 ( Lispinus ; Guinea; Ivory Coast).

- DISTRIBUTION: Guinea, Ivory Coast, Togo, Cameroon, Gabon, Tanzania. acutepunctatus Bernhauer, 1929 , see: bernhauerianus Scheerpeltz, 1933 . acutepunctatus Bernhauer, 1929 i: 344 ( Lispinus ; Type locality: Luzon: Haight’s Place, Balbalan).

— Scheerpeltz, 1933: 1012 ( Lispinus ; catalog).

- DISTRIBUTION: Philippines.

aeneiceps Cameron, 1925b: 236 , 240 ( Lispinus ; Type locality: Penang).

— Scheerpeltz, 1933: 1012 ( Lispinus ; catalog).

— Cameron, 1936: 4 ( Lispinus ; Society Islands).

— Cameron, 1936a: 29 ( Lispinus ; Java).

— Coiffait, 1976: 220 ( Lispinus ; Tahiti).

- DISTRIBUTION: Malaysia, Indonesia, Society Islands.

aequalis Fauvel, 1878f: 205 ( Lispinus ; Type locality: Nouvelle-Guinée, Ansus, Ramoi; Iles Key).

— Bernhauer and Schubert, 1910: 21 ( Lispinus ; catalog).

— Bernhauer, 1922b: 166 ( Lispinus ; Seychelles).

— Cameron, 1931: 356 ( Lispinus ; New Guinea).

— Cameron, 1937a: 87 ( Lispinus ; New Guinea).

- DISTRIBUTION: New Guinea, Indonesia, Seychelles.

aequicollis Bernhauer, 1929 i: 350 ( Lispinus ; Type locality: Surigao, Dapa).

— Scheerpeltz, 1933: 1012 ( Lispinus ; catalog).

- DISTRIBUTION: Philippines.

aequipunctatus LeConte, 1868: 50 ( Lispinus ; Type locality: Cañada de los Diegos , New Mexico).

— LeConte, 1877: 250 ( Lispinus ; characters).

— Fauvel, 1878d: 177 [= 1878e: 13] ( Lispinus ; characters; New Mexico).

— Bernhauer and Schubert, 1910: 21 ( Lispinus ; catalog).

- DISTRIBUTION: USA.

aethiops Eppelsheim, 1895 , see: Nacaeus .

affinis Cameron, 1943a: 336 ( Lispinus ; Type locality: New Guinea: Moroko).

- DISTRIBUTION: New Guinea.

africanus Bernhauer, 1929 , see: Nacaeus .

agnatus Fauvel, 1904e: 297 ( Lispinus ; Type locality: Tananarive; forêt de Moramanga). — Bernhauer and Schubert, 1910: 21 ( Lispinus ; catalog).

- DISTRIBUTION: Madagascar.

alutaceipennis Scheerpeltz, 1933: 1012 ( Lispinus ; replacement name for alutipennis Bernhauer ).

- DISTRIBUTION: Philippines.

alutipennis Bernhauer, 1929 i: 354 ( Lispinus ; [preoccupied]; Type locality: Palawan, Binaluan ).

— Scheerpeltz, 1933: 1012 ( Lispinus ; synonym of alutaceipennis ).

alutaceipennis Bernhauer, 1937 , see: kenyanus Herman, 2001 .

alutaceus Fauvel, 1878f: 205 ( Lispinus ; Type locality: Nouvelle-Guinée, Hatam).

— Bernhauer and Schubert, 1910: 21 ( Lispinus ; catalog).

— Bernhauer, 1926f: 312 ( Lispinus ; Buru).

- DISTRIBUTION: New Guinea, Indonesia.

alutipennis Bernhauer, 1921b: 66 ( Lispinus ; Type locality: Bolivien: Yuracaris ).

— Scheerpeltz, 1933: 1012 ( Lispinus ; catalog).

- DISTRIBUTION: Bolivia.