Tinodes ivanovi, Gibon, 2017

Gibon, François-Marie, 2017, New species of Tinodes from Madagascar (Trichoptera, Psychomyiidae), Zootaxa 4318 (2), pp. 364-376 : 365

Genus Tinodes Curtis 1834

In Madagascar, two groups of closely related Tinodes species can be identified; besides these a single species is currently incertae sedis. The T. isalo Species Group is characterized by the width of the phallic apparatus and the presence of two internal sclerites: a preapical one, which is variable or missing and an apical one, which is stout, short and spine-shaped. The intermediate appendage branches are moderately long, shorter than the preanal appendages; they are curved ventrad, finger-shaped, wider apically, and with a group of preapical spines. It includes Tinodes isalo Melnitsky & Ivanov 2016 , Tinodes ivanovi n. sp., Tinodes tafo n. sp., and Tinodes helix n. sp. Tinodes mohelia Johanson & Oláh 2007 , from the Comoros, is related to this group. In the T. irwini Species Group, the phallic apparatus is not as wide as in the T. isalo Species Group. The intermediate appendage branches are short or moderately long, their basal part is straight or weakly curved ventrad and their distal part is enlarged, curved dorsad or dorsally developed and strongly modified, generally with spines. This T. irwini Species Group includes Tinodes irwini Johanson & Oláh 2007 , Tinodes schlingeri Johanson & Oláh 2007 , Tinodes dehadawate Johanson & Oláh 2007 , Tinodes harawa Johanson & Oláh 2007 , Tinodes forcipatus n. sp., and Tinodes marcossi n. sp. These Malagasy groups are distinguished by the width of their phallic apparatus from the continental African species, most of the latter having the phallic apparatus gently curved, long, and slender. Another interesting fact is the high proportion of Malagasy species with asymmetrical intermediate appendage branches, as noted by Johanson & Oláh (2007). Only two Malagasy species have symmetrical intermediate appendage branches: T. isalo in the T. isalo Group and T. forcipatus n. sp. in the T. irwini Group.