Tipula (Sivatipula) pseudofilicornis, Men, 2020

Men, Qiulei, 2020, Three new species of the subgenus Tipula (Sivatipula) from Yunnan and Tibet, China (Diptera: Tipulidae), with an updated key for Chinese species, Zoological Systematics 45 (1), pp. 59-68 : 63-65

Tipula (Sivatipula) pseudofilicornis sp. nov. ( Figs 16–29 )

Diagnosis. Body brownish-yellow in general; thorax dark brown with three brownish-yellow stripes on prescutum, the median one divided by brown vitta; antenna longer than the body length; wing brownish-yellow with a light brown stigma; abdomen brownish-yellow with lateral stripes brown, segments six to eight entirely black, hypopygium black; tergite nine with a pair of horn-shaped processes on lateral corners and a broad median lobe; inner gonostylus with an obtuse lobe and a finger-shaped process on mesal region of outer surface.

Material examined. Holotype male, China, Tibet, Linzhi, Chayu , Shangchayu , 28°43'N, 96°46'E, 8 Jun 2019, leg. Qiulei Men. Paratypes. 3 males, same data as holotype .

Description. Adult length (male, n =4): body 13.1–13.3mm (not including antenna), wing 15.7–15.9 mm, antenna 15.0– 15.2 mm.

Head. Brownish-yellow with rostrum dark brown laterally ( Fig. 16 ). Nasus brown with black setae apically. Vertex grayish-brown, lacking of marking ( Fig. 16 ). Head including vertex and rostrum densely covered with black setae. Antenna 12-segmented, significantly longer than body; scape, pedicel and first flagellomere brownish-yellow, rest flagellomeres yellowish-brown, each flagellomere subequal in length, cylindrical, with black verticils. Palpus dark brown.

Thorax. Pronotum brown ( Fig. 17 ). Prescutum brown with three brownish-yellow stripes, the median one divided by brown vitta ( Fig. 18 ). Scutum dark brown with two black marks, scutellum brown, postnotum brown with a grayish median vitta ( Fig. 18 ). Pleuron entirely brownish-yellow ( Fig. 17 ). Legs very slender, coxae and trochanters brownish-yellow, femora brownish-yellow with a darker apex, tibiae and tarsi dark brown. Halter with stem yellow, knob black. Wing brownish-yellow, cell sc darker than ground color, stigma light brown ( Fig. 19 ). Rs distinctly longer than R 3, petiole of cell m 1 subequal in length to m-m, 1/3 the length of discal cell ( Fig. 19 ).

Abdomen. Segments one to five brownish-yellow with black lateral stripes, segments six to eight wholly black. Hypopygium black ( Fig. 20 ). Tergite nine with a pair of horn-shaped processes on lateral corners, median lobe broad, shallowly concaved medially, the flange between horn-shaped processes and median lobe deeply concaved ( Figs 21–23 ). Outer gonostylus with apical margin and anterior margin forming a right angle, posterior margin terminagting in a small lobe ( Fig. 24 ). Inner gonostylus broad basally, protruding into a beak, a small obtuse lobe and a finger-shaped processes from its mesal region of outer surface ( Figs 25–26 ). Sternite nine possessing a horn-shaped process which is densely covered with long setae ( Fig. 27 ).

Semen pump with compressor apodeme V-shaped, with a pair of ridges on its arms ( Fig. 28 ). Posterior immovable apodeme being one median arm which curved dorsally. Anterior immovable apodeme flattened, expanded in lateral view ( Fig. 29 ). Aedeagus narrow, elongated, tubular, more than 10.0 times longer than the length of semen pump.

Distribution. China (Tibet).

Remarks. The new species is closely allied to T. (S.) filicornis Brunetti, 1918 , another Indian species by similar structure of the hypopygium. It can be easily separated from T. (S.) filicornis by the outer gonostylus which forms a right angle on dorsal margin. T. (S.) filicornis has its outer gonostylus Y-shaped and lacking of such right angle.

Etymology. The specific epithet is the name of its related species “ filicornis ” with a Latin prefix “ pseudo ”, emphasizing the morphological similarity of the new species and its related species.

Figures

Figures 16–20. Tipula (Sivatipula) pseudofilicornis sp. nov. 16. Head, dorsal view. 17. Thorax, lateral view. 18. Thorax, dorsal view. 19. Wing. 20. Hypopygium, lateral view.

Figures 21–29. Tipula (Sivatipula) pseudofilicornis sp. nov. 21. Hypopygium, lateral view. 22. Tergite nine, dorsal view. 23. Tergite nine, posteroventral view. 24. Outer gonostylus, inner view. 25. Inner gonostylus, outer view. 26. Inner gonostylus, inner view. 27. Median processes on sternite nine, dorsal view. 28. Compressor apodeme, dorsal view. 29. Semen pump, lateral view.