Tipula (Vestiplex) medialobata, Men & Sun & Starkevich, 2023
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publication ID |
https://doi.org/ 10.5852/ejt.2023.856.2037 |
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publication LSID |
lsid:zoobank.org:pub:50E202D5-BB02-469B-B946-27736CA86162 |
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DOI |
https://doi.org/10.5281/zenodo.7626058 |
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persistent identifier |
https://treatment.plazi.org/id/769474A5-416C-4C8A-931C-002CB756090D |
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taxon LSID |
lsid:zoobank.org:act:769474A5-416C-4C8A-931C-002CB756090D |
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treatment provided by |
Felipe |
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scientific name |
Tipula (Vestiplex) medialobata |
| status |
sp. nov. |
Tipula (Vestiplex) medialobata sp. nov.
urn:lsid:zoobank.org:act:769474A5-416C-4C8A-931C-002CB756090D
Figs 16–27 View Figs 16–20 View Figs 21–27
Diagnosis
Tipula (V.) medialobata sp. nov. can be recognized by the tergite nine having posterolateral corners ear-like and black narrow, serrated rods; gonocoxite having two small points on apex and a black long spine on mesoventral portion; clasper of gonostylus with medial surface raised into blackened, nearly rectangular lobe and long, blade-shaped dorsal lobe of A9S apically blackened and sharp; antenna reaching base of wing if bent backwards, flagellum brown; wing light brown with dark brown stigma.
Etymology
The specific epithet is derived from the Latin ‘ lobata ’ with prefix ‘ media ’, referring to the lobe in the middle area of the clasper of the gonostylus.
Material examined
Holotype CHINA • ♂; Yunnan, Gongshan County; Bingzhongluo, Zhanatong ; 11 Jul. 2019; Q. Men and Lin Lv leg.; AQNU.
Paratypes CHINA • 3 ♂♂; Yunnan, Gongshan County; Bingzhongluo, Gongdangshenshan Mountain ; 12 Jul. 2019; Q. Men and Lin Lv leg.; AQNU .
Comparative material examined
INDIA • 3 ♂♂; holotype of Tipula (Vestiplex) tillyardana Alexander, 1970 ; Assam [Arunachal Pradesh]; Shergaon, Kameng; North East Frontier Agency ; alt. 6100 feet; 5 May 1961; F. Schmid leg.; antenna, leg, wing and genitalia on slide mounted; USNM .
Description
Male ( Figs 16–27 View Figs 16–20 View Figs 21–27 )
MEASUREMENTS. Body length 12.5–12.7 mm (excluding antenna, n = 4), wing length 18.1–18.3 mm (n = 4), length of antenna 3.6–3.7 mm (n = 4). General body coloration yellowish-brown.
HEAD ( Fig. 16 View Figs 16–20 ). Rostrum with nasus yellowish, densely covered with brown setae. Eye black. Occiput and vertex yellowish, medially with dark-brown stripe broadened medially and narrowed at each end ( Fig. 16 View Figs 16–20 ). Antenna 13-segmented, if bent backward reaching base of wing. Scape yellow, cylindrical, slightly expanded at apex. Pedicel yellow, oval. First flagellomere yellowish-brown, rest flagellomeres brown, basal four flagellomeres almost same length to each other, remaining segments progressively shortened and thinned, last flagellomere oval. Each flagellomere except first one enlarged at base, with five longest yellow verticils. Verticils slightly longer than corresponding flagellomeres, surface of each flagellomere densely covered with short brown setae. Palpus yellow.
THORAX ( Figs 17–18 View Figs 16–20 ). Pronotum yellow with lateral regions darker in coloration, medially with dark spot. Prescutum and presutural scutum yellowish-brown, with four brown longitudinal stripes, narrowly bordered by darker brown ( Fig. 17 View Figs 16–20 ). Intermediate pair separated. Interspaces thinly dusted with gray, densely covered with short yellow setae ( Fig. 17 View Figs 16–20 ). Postsutural scutum brown, scutal lobes each with two darker brown separated triangular spots. Scutellum brownish with dark brown median vitta, densely covered with yellow setae ( Fig. 17 View Figs 16–20 ). Mediotergite dark brown with broad black median line ( Fig. 17 View Figs 16–20 ). Pleura brownish-yellow, thinly dusted with grey ( Fig. 18 View Figs 16–20 ). Leg with coxa and trochanter yellow, femur yellow with darkened tip, tibia and tarsal segments dark brown. Wing light brown, cell c more yellowish than ground color, stigma dark brown, variegated by light areas including apical area of cell c, and basal half of 1 st cell of cell r 1, light band across apical area of cell r 1, base of cells r 3, r 4 and r 5, and medial area of discal cell; rest light area including basal and apical areas of basal radial cell, distal and basal areas of bm, and apical area of posterior cubital cell ( Fig. 19 View Figs 16–20 ).
WING VENATION. R 1 complete, R 3 1.2 × as long as R 2+3+4, R 4 1.1 × as long as Rs, R 5 curved at apical half, base of R 5 distinctly longer than r-m, discal cell narrow, at least 2.3 × as long as petiole of cell m 1, M 1 almost 4.4 × as long as its petiole ( Fig. 19 View Figs 16–20 ). Halter pale yellow, knob basally black, densely covered with black setae, apically pale yellow.
ABDOMEN ( Fig. 20 View Figs 16–20 ). Basal four tergites yellow with brown lateral and intermittent median stripes, tergites 5–8 brown with yellow lateral regions, basal five sternites entirely yellow, sternites six to eight gradually heavier to dark brown, hypopygium dark brown ( Fig. 20 View Figs 16–20 ).
HYPOPYGIUM ( Figs 21–27 View Figs 21–27 ). Hypopygium with tergite nine and sternite nine separated except base ( Figs 21–22 View Figs 21–27 ). Tergite nine completely divided at midline by pale membrane ( Fig. 21 View Figs 21–27 ). Dorsal portion of tergite nine shallowly emarginated medially, terminating into pair of ear-like lobes on lateral corners which is densely covered with setae ( Fig. 21 View Figs 21–27 ). Ventral portion of tergite nine possessing pair of black narrow rods which are serrated on inner margins ( Fig. 21 View Figs 21–27 ). Gonocoxite separated from sternite nine except ventral area, massive, with two small points on apex and black long spine on mesoventral portion only observed in ventral view ( Figs 22–23 View Figs 21–27 ). Lobe of gonostylus narrow and elongated ( Fig. 24 View Figs 21–27 ). Clasper of gonostylus with upper beak broad and obtuse, preceded by black ridge along mesal margin; lower beak blackened oval lobe; medial surface raised into blackened, nearly rectangular lobe ( Fig. 24 View Figs 21–27 ). Sternite nine with ventral lobe of A9S thumb-like, dorsal lobe of A9S blade-shaped, broad at base, gradually narrowed to sharp black apex ( Fig. 23 View Figs 21–27 ). Adminiculum relatively short, nearly triangular in ventral view, truncated apically; median portion protruded into distinct sclerite ( Fig. 23 View Figs 21–27 ). Semen pump with central vesicle swollen ( Fig. 26 View Figs 21–27 ). Ejaculatory apodeme with deep V-shaped median incision, lateral arms expanded apically ( Fig. 25 View Figs 21–27 ). Posterior immovable apodeme narrow ( Fig. 26 View Figs 21–27 ). Anterior immovable apodeme distinctly broad, round in dorsal view ( Fig. 27 View Figs 21–27 ). Aedeagus thick at base, gradually narrowed to apex, more than 3.5 × as long as semen pump ( Fig. 26 View Figs 21–27 ).
Female
Unknown.
Remarks
Tipula (V.) medialobata sp. nov. is assigned to the T. (V.) subticta species group. The new species is very similar to T. (V.) tillyardana Alexander, 1970 , from Arunachal Pradesh, India, by details of tergite nine. Posterolateral corners of tergite nine are extended into lobes in both species, but still differs by the shape: T. (V.) medialobata sp. nov. has lobes ear-shaped while the lobes are obliquely truncated in T. (V.) tylliardana ( Alexander 1970: fig. 37). They can also be separated by the gonocoxite which apically has two points in the new species while only a small blackened spine is present in T. (V.) tylliardana ; the mesoventral portion of the gonocoxite has a black spine in the new species while a curved arm with microscopic points is present in T. (V.) tylliardana . Also, the new species is distinguished by a clasper of the gonostylus bearing a blackened lobe on the medial surface while such a lobe is absent in T. (V.) tylliardana .
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.