Toccolus javanensis

Kury, Adriano B., 2008, On the systematic position of Dino Loman and Toccolus Roewer (Opiliones, Laniatores, Epedanidae), with the description of a new species from western Java, Indonesia, Zootaxa 1932, pp. 61-68 : 63-65

Toccolus javanensis sp. nova

( Figs 1–8 )

Type material. AMNH AK 172 ɗ holotype “Poentjak, S.F. 1920” [ Indonesia, Java Island, Jawa Barat Province, Bogor Regency, Gunung Puncak, -6.70585° 106.976°]. WWF Ecoregion: IM0167 (Western Java Montane Rain Forests).

Etymology. From the island of Java, to contrast with the other species, known from continental SE Asia.

Description of male holotype.

Measurements. ID 0.9, CL 1.9, AL 1.0, CW 2.4, AW 2.7. See Table 1 for measures of appendages.

Dorsum ( Figs 1–2 ). Peltidium subrectangular, with sides concave. Cheliceral sockets very deep. Ocularium very low, ovoid, with a median small tubercle. Three ill-defined mesotergal areas, area I and II fused together. Peltidium and all free tergites smooth and unarmed.

Chelicera ( Figs 1–3 ). Basichelicerite very robust, two-thirds as long as peltidium, with bulla attenuate, armed with dorsal, dorso-ectal and dorso-mesal strong acuminate tubercles. Cheliceral hand swollen (presumably only in male). Dentition of fingers composed of (1) basal larger isolated teeth and (2) distal smaller grouped teeth. Teeth of movable finger in double row, forming a locking area for fixed finger.

Pedipalpus ( Figs 1–2, 4–5 ). Coxa very long, with 2 strong dentiform dorsal apophyses, anterior one very large, anchor-shaped and ventro ectal subdistal setiferous tubercle. Trochanter short, with two robust ventral divergent setiferous tubercles and one dorsal setiferous tubercle. Femur straight, a little shorter than peltidium, armed dorsally with two rows of small tubercles and ventrally with a row of 6 large spines. Femur mesal subdistal with two large spines. Patella ventro-ectally with 1 and ventro-mesally with 2 large setiferous tubercles. Tibia ventro-ectally with 4 and ventro-mesally with 3; tarsus ventro-ectal and ventro-distal with 4 large setiferous tubercles each. Both tibia and tarsus flattened dorsally, neither bulbous nor forming a subchela with femur.

Legs. All podomeres slender and unarmed. Coxa IV very small, reaching only posterior border of area I/ II, as is typical in epedanids (contrasting to larger coxa, almost reaching end of mesotergum, as for example in Toccolus minimus , based on Roewer’s figure), distal part visible in dorsal view, surpassing peltidium at posterior line of carapace. Tarsal counts: 8(2)/21(3)/?/10.

Color. Body and appendages uniform dark orange brown with faint black reticulation. Posterior margin of combined areas I–II with a black rim.

Genitalia ( Figs 6–8 ). Pars distalis of truncus with a broad excavation ventrally forming a thin medial lamina, which bends back upon itself by means of left and right hyaline folds, leaving a long V-shaped opening in the middle. [the previous sentence is not clear. A thin median lamina is not evident in the figure. The truncus seems to have more of an excavation, rather than being concave. Also, the line in the lateral view (f.7) seems to be a sharp edge and so should also be visible in the ventral view. The dorsal part of pars distalis slopes gently, forming a terrace from which the glans projects. The numerous paired setae are distributed as follows: 1) 1 pair ventro-basal on the pars distalis of truncus; 2) 1 pair ventral strongly asymmetrical on the base of the folds; 3) 3 ventral pairs along the hyaline part of the folds; 4) 5–6 strongly curved setae on each side encircling the glans insertion; 5) 1 pair of erect setae flanking the glans. Glans formed by a follis including two stages, basalmost column more or less rigid, and a complex of haematodochae surrounding the stylus, which is not free.

Figures

FIGURE 1. Toccolus javanensis sp. nova, male holotype from Gunung Puncak, Java, habitus, dorsal view. Scale bar = 1 mm.

FIGURES 2 – 5. Toccolus javanensis sp. nova, male holotype from Gunung Puncak, Java. 2, habitus, lateral view; 3, Right chelicera, frontal view; 4, Left pedipalpal trochanter, ectal view, detail; 5, Right pedipalpal patella to tarsus, ventral view. Scale bars = 1 mm.

FIGURES 6 – 8. Toccolus javanensis sp. nova, male holotype from Gunung Puncak, Java, penis distal part. 6, dorsal view; 7, lateral view; 8, ventral view. Scale bar = 0.1 mm.

Tables

TABLE 1. Appendage measurements of Toccolus javanensis. Abbreviations: claw, pedipalpal claw; Fe, femur; Mt, metatarsus; Pa, patella; Ta, tarsus; Ti, tibia; Tr, trochanter.

	Tr	Fe	Pa	Ti	Mt	Ta	claw
Pedipalpus	0.6	2.6	4.1	1.5	–	1.3	1.0
Leg I	0.3	1.7	0.6	1.4	2.3	1.4	–
Leg II	0.5	2.3	0.8	?	?	?	–
Leg III	0.5	2.2	0.8	1.7	2.7	?	–
Leg IV	0.5	2.9	0.8	2.0	3.5	1.8	–