Trichophallus uniformis ( Willemse, 1966 ) Ingrisch, 2024

Ingrisch, Sigfrid, 2024, Revision of the genus Trichophallus Ingrisch, 1998 with notes on the genera Secsiva Walker, 1869 and Subrioides C. Willemse, 1966 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini), Zootaxa 5442 (1), pp. 1-66 : 58-59

Trichophallus uniformis ( Willemse, 1966) comb. nov.

Figs. 30–31 : 110155F + 110155M

Subria uniformis C. Willemse 1966 , Publ. natuurh. Gen. Limburg 16: 10, fig. 18.1966 Willemse, Publ. Natuurhist. Genoot. Limburg 16: 10—1979 Nishida, Pacific Insects 20: 27–32 >> Type data—Otte, D. 1997. Orthoptera Species File 7:70 >> Note: 1996 Ingrisch pers. comm.: probably not Subria >> Subria uniformis

Holotype (male), allotype (female), New Britain, Gazelle Penins. Bainings St. Pauls 350m, 07.09.1955, J.L. Gressitt ( BPBM Honolulu Museum), both not re-exemined.

Other specimens studied: Papua New Guinea: New Britain, Gazelle Pen., Bainings: St. Paul‘s, elev. 350 m, 7 September 1955 (coll. J.L. Gressit)— 1 male, 1 female ( Bishop Museum , Honolulu ( BPBM)); New Britain, Gazelle Pen., Bainings: St. Paul‘s , elev. 350 m, 9 September 1955 (coll. J.L. Gressit)— 1 female (Bishop Museum, Honolulu ( BPBM)); New Britain, Gazelle Pen., Gaulim, elev. 130 m, 28 October 1962 (coll. J. Sedlacek)— 1 male (Bishop Museum, Honolulu ( BPBM)); New Britain, Gazelle Pen., Mt. Sinewit , elev. 1100–1200 m, 15–16 November 1962 (coll. J. Sedlacek)— 1 male (Bishop Museum, Honolulu ( BPBM)); New Britain, Gazelle Pen., Talliligap, elev. 300 m, 17–18 December 1962 (coll. J. Sedlacek)— 1 female (Bishop Museum, Honolulu ( BPBM)); New Britain, Gazelle Pen., Upper Warangol , elev. 250–700 m, 6–7 December 1962 (coll. J. Sedlacek)— 1 male (Bishop Museum, Honolulu ( BPBM)); New Britain, Kerawat, elev. 135 m, 20–25 November 1959 (coll. T.C. Maa)— 1 male (Bishop Museum, Honolulu ( BPBM)); New Guinea ( NE), East New Britain, „New Guinea “, Rabaul , 22 October 1932 (coll. J.L. Froggatt)— 1 female ( NHM London) .

Records not examined: West New Britain, New Britain, Linga Linga Plain, W. of Willaumoz Pen. l m., 12 April 1956 (coll. J.L. Gressitt)— 1 male paratype (Naturalis Leiden) .

Diagnosis. Males of T. uniformis can be recognised by the shape of the elongate cerci that carry a hump with a spine near base, the subgenital plate is of medium length and the last abdominal tergite provided with a short apical excision; the male cerci have the reduced acute internal process in apical area; titillators are provided with one pair of main sclerites. Females have the subgenital plate projecting more laterally than in other species of the genus and the apical projections are markedly curved and reaching about mid-length of the conical projections of the ventral ovipositor valves.

Description. Medium sized to large species. Tegmina reaching apical third of hind tibia. Femora with the following number of spines on ventral margins: fore femur 4–7 external, 3–6 internal; mid femur 6–7 external, 2–4 internal near base; hind femur 9–12 external, 11–15 internal.

Male. Stridulatory file with about 130–136 teeth. Tenth abdominal tergite terminating into two short projections, with a wide and concave or truncate excision in between. Epiproct rectangular with a deep pit near base, apex convex or truncate. Paraprocts with internal margin swollen and slightly projecting. Cerci long and narrow, conical and slightly sinusoidal curved; with a large, acute, internal projection before middle of length; apex of cercus often collapsed in set specimens, with or without a spinule at tip. Subgenital plate setose, roundly excised at apex; styli long and thin, of nearly equal length with excised area.

Titillators short and stout, apex covered with a rounded sheath; with a bilobate bursa at each side laterally of titillator apex; ventral-medial lobe conical, granular, little sclerotised, margin spinulose; dorsal-proximal lobe large, semi-ovoid behind apex of titillators, and at bottom of internal surface provided with a row of large bristles surpassing apex of lobe; at dorsal-proximal side lobe conically projecting with rounded apex; external surface of bursae with a distinctly granular and slightly sclerotised area.

Female. Tenth abdominal tergite divided in midline. Subgenital plate with entire ventral area short, transverse, grooved, and with an apical carina that is slightly curved backwards on both sides towards middle, divided in midline; projections of ventral area compressed, almost straight but pointing apical-medial, with ventral-medial margin concave and apex obliquely truncate or convex; lateral areas divided by a carina in about middle of height and a furrow dorsal of carina, area dorsal of furrow bent lateral. Eight abdominal tergite with lateral-posterior angle excised; a lobe of the intersegmental membrane that is carrying the stigma is stiffened and swollen and partly covering the lateral area of the subgenital plate. Dorsal margin of ventral ovipositor valves at base with an elongate, swollen, setose, oblique triangular projection with rounded angle.

Variations: (1) Lo003F001: Subgenital plate lateral area with strong 90° fold in middle; just ventral of fold with a high lamella; ventral and ventral apical margins swollen; dorsal margin little sinusoidal.—(2) Lo003F002: Tenth abdominal tergite divided in midline. Epiproct triangular; dorsal surface grooved. Cerci narrow, slightly curved; apex pointed. Subgenital plate in short basal part with surface slightly grooved; afterwards divided into two widely spaced, compressed, narrow lobes approaching backwards; lateral areas bent from ventral surface in greater than 90° angle, strongly concave with swollen margin and weak carina in middle. Ovipositor falcate; dorsal margin of ventral valves with conical, hairy appendage at base.

Coloration. Uniformly yellowish brown. Pronotum with faint indication of brown medial band split by a white line. Tegmen nearly transparent with scattered brown dots and brown dorsal margin.

Measurements.—body: male 22–27, female 19–28; pronotum: male 5.6–6.2, female 5.5–6.5; tegmen: male 33– 38, female 34.0–38.5; hind femur: male 16.5–18.0, female 13–18; antenna: male 60–75, female 65–85; ovipositor: 11.5–12 mm.

Figures

FIGURE 30. Trichophallus uniformis (Willemse, 1966) male: A, stridulatory file; B, habitus male with left wings spread; C, subgenital plate and cerci in ventral view; D, last abdominal tergite and cerci in dorsal view; E–G, titillators in ventro-apical (E), apical (F), and dorsal-proximal view (G). Male from Linga Linga Peninsula (New Britain).

FIGURE 31. Trichophallus uniformis (Willemse, 1966) females:A, E–F, subgenital plate and dorsal lobes of ventral ovipositor valves in lateral view; B, subgenital plate in ventral view; C, abdominal apex in lateral view; D, ovipositor.—A–D, female from Gazelle Peninsula; E, female from Rabauel; F, female from Vagau.—Abbreviations: de, basal dorsal expansion of subgenital plate; pl, plate with apical prolongations; op, oval projection from dorsal margin of ventral ovipositor valve.