Triptognathus uniguttatus (GRAVENHORST), 1978

A. M, 2011, Illustrated key to the genera of the subtribe Amblytelina of Palaearctic (Hymenoptera, Ichneumonidae, Ichneumoninae, Ichneumonini), Linzer biologische Beiträge 43 (1), pp. 597-711 : 682-684

Triptognathus uniguttatus (GRAVENHORST) (Plate 16)

Ichneumon uniguttatus GRAVENHORST 1829 - Ichneumonologia europaea 1: 310,,. Triptognathus uniguttatus: HEINRICH 1978 - East. Pal. Ichn.: 60-61. M a t e r i a l e x a m e n e d:: Kokchetav, Akmol. obl., Karavaev, 01.08.190 8.: okr.

Antonovki, Pishpekskogo u. [Kirgizija], P. Arkhangel'skij, 19.05.192 2.: Turcja, Ispir, 2500

m, 15.07.197 4, leg. T. Huflejt, " Triptognathus atripes Grav. (?)", det. A. Rasnitsyn.

F e m a l e

F l a g e l l u m: Bristle-shaped, with 54 segments, moderately thick, without white annulus, only just widened and very slightly flattened ventrally beyond middle; segments slightly elongated, first segment short, only 1,7 times longer than the width at apex, already segment 4 square from lateral; segments of base sharply separated (differentiated), other segments more or less merged. Flagellum 1,3 times shorter than the front wing and 1,5 times shorter than body length.

H e a d: Head contour from front quite slightly narrowed downwards, only just transverse, only 1,1 times wider than height; genae from front (malar space) 4,3 times shorter than height of eye (0,2), genae visible from front on 0,3 from base of an eye; head contour from above transverse, 2,2 times wider than length in middle and 1,8 times along an external contour, almost linearly narrowed backwards behind eyes. Vertex from lateral linearly sloping from hind border of ocelli to occipital carina; temples long, in middle 1,5 times longer than longitudinal diameter of an eye, below middle parallel to hind margin of an eye; occipital carina sharp all round, from above sharp and high elevated (but not so much as in Triptognathops HEINRICH ), uniformly roundly impressed, far not reach level of eyes and hind ocelli; hypostomal carina visible from lateral to 0,2-0,3 height of an eye, merged with occipital carina before base of mandible; abscissula rather high, equal 0,7 of mandible base width; malar space equal to mandible base width; mandibles rather narrow, evenly narrowed from base to apex, with more or less sharpened apex, without lower tooth; clypeus convex at base, 2,2 times wider than length, with straight thinned front margin and rounded lateral corners, separated from face by deep, broad impression; clypeal foveae small, deep; labrum equal by breadth to front margin of clypeus, rounded, rather long; middle field of face slightly separated, short, rather sharply elevated, in middle approximately equal by breadth to lateral fields; antennal cavities short, rather deeply impressed, reach borders of eyes and far not reach front ocellus level, without lateral tubercles and with strong tooth between antennal fossae; margins of antennal fossae slightly elevated above face surface; ocelli of normal size, diameter of lateral ocellus 1,4 times less than distance from ocellus to eye; ocellar triangle slightly elevated. Surface of clypeus with big deep punctures; middle and lateral fields densely punctured to wrinkly-punctured on theirs borders; frons roughly wrinkly-punctured to sharplywrinkled at area of front ocellus; temples smoothly wrinkly-punctured; surface of head without microsculpture.

T h o r a x: Collar of pronotum from above long with rounded front margin; transverse furrow of pronotum deep, narrow; pronotal ridge not swollen; epomiae sharp, high; pronotal base gradually curved, slightly sinuous. Mesonotum moderately convex, only just (1,1 times) longer than width; notauli absent; surface of mesonotum densely punctured, without microsculpture; axillary tongue weak but distinct; subalarum high, moderately thick, not sharpened; speculum densely wrinkly-punctured, same as other part of upper third of mesopleurae; mesopleural fovea sharply expressed, deep; mesopleural suture straight, deep, interrupted by sharp ribs; mesopleurae at bottom with bend from lower third; sternauli sharp, triangular, up to hind third of mesopleurae; surface of mesopleurae on the larger part roughly wrinkly-punctured, without microsculpture; scutellum from lateral moderately elevated above postscutellum, from above almost flat, laterally not carinated, its surface densely punctured by large oblong punctures. Hind margin of metanotum with broad triangular projections opposite of lateral longitudinal carinae. Length of horizontal part of propodeum 1,5 times less than length of area posteromedia in middle; carinae of propodeum with exception of costulae expressed; basal area short and deep; area superomedia rectangular, from square to slightly transverse, 1,2 times wider than length; carinae, enclosing areae dentiparae and areae spiraculifera high and thus form at apices of areae dentiparae lamellar broadening like tooth (but not a tooth); spiracles long, slit-shaped, along external contour 4 times longer than width. Surface of propodeum, including metapleurae, roughly-wrinkled, without microsculpture.

L e g s: Rather slender, hind tarsi long and slender. Middle tibiae with more strong spinules comparatively tibiae 1 and 3. Claws rather long, uniformly curved.

W i n g s: Areolet pentagonal, practically symmetrical with broad base, external vein of apex shorter than internal one; stigma short, broad, more or less light; radial cell broad; radius almost straight; nervulus interstitial; ramulus long; all veins dark; membrane of wing slightly darkened. Front wing longer than length of flagellum and shorter than body length.

A b d o m e n: Amblypygous, from above broadly-oval, apical tergites not hidden, sheath of ovipositor from above not protrude; second tergite transverse, at apex 1,5 times wider than length. First tergite from lateral with rather sharp bend between petiolus and postpetiolus, from lateral sharply bordered by carinae and with strong transversal ribs between them; from above petiolus sharply broadened to rectangular behind spiracles postpetiolus; distance between spiracles 1,5 times more than distance from spiracles to hind margin of tergite; middle field of postpetiolus sharply expressed and high elevated, 1,5 times wider than lateral fields, its surface roughly longitudinally striated, lateral fields also with longitudinal wrinkles. Gastrocoeli small, only just impressed, short; thyridia weak, short, many times narrower than interval between them (only just marked); lunulae only just expressed, small; interval between gastrocoeli with very slight longitudinal striation, surface of second and third tergites very densely and indistinctly punctured, with slight shine. Hypopygium more or less flattened, with membranous hind margin and with fascicle of long stiff bristles at apex (as in Eutanyacra CAMERON ); sternites 4-7 with short delicate pubescence; only sternites 2-3 with longitudinal fold.

C o l o r a t i o n: Body black, small spots on vertex and tergites 2-3 of abdomen red.

S i z e: Body length: 12,3; front wing: 10,5; flagellum: 8,0 mm.

M a l e

Flagellum of male with 43 segments, with dark bacilliform tyloides on segments 6-17, segments of flagellum not ribbed, densely merged. Mandibles narrow, without lower tooth. Hind margin of collar of pronotum high, sharp, transverse furrow deep. Sculpture of mesopleurae less rough than of females; horizontal surface of scutellum convex. Transverse carinae at apices of areae dentiparae remind a tooth in a greater extent than in females. Hypopygium at apex with long narrow process. Collar of pronotum, spots in corners of pronotum, subalarum, tegulae, spots at base of front wings and scutellum white; coloration of abdomen is the same as of females; tibiae and tarsi of front and middle legs, femora of front completely, of middle one on the larger part reddish-black with yellow.

N o t e s: The male available in our disposal, determined by A. RASNYTSIN as Triptognathus ? atripes (GRAV.) (RASNITSYN 1981) has occipital carina considerably elevated at horizontal part as in Triptognathops HEINRICH , practically not expressed tyloides and considerably more rough sculpture of a body. It appreciably differs from the described above and identified by G. HEINRICH (see HEINRICH 1978) as uniguttatus male. H. TOWNES (1965) has synonymized Triptognathus atripes and T. uniguttatus , considering of white scutellum, white spot on subalarum and a median apical white mark on tergites 6 and 7 presence in T. atripes female as insignificant differences. The said calls into question the synonymization of Triptognathus atripes and T. uniguttatus .

For this reason, we leave for Triptognathus uniguttatus (GRAV.) the former name under what it has been identified by G. HEINRICH.