Ungla quchapampa Tauber

Tauber, Catherine A., Sosa, Francisco, Albuquerque, Gilberto S. & Tauber, Maurice J., 2017, Revision of the Neotropical green lacewing genus Ungla (Neuroptera, Chrysopidae), ZooKeys 674, pp. 1-188 : 65-67

Ungla quchapampa Tauber sp. n. Figs 77, 78, 79, 80, 144e

Holotype

(Figs 77, 79, 80, 144e). Male (MCZ). Bolivia, Cochabamba, Chapare, Steinbach.

Etymology.

The type locality of this species is the Andean valley of Cochabamba in central Bolivia. The locality is also known by its Quechua name - Quchapampa. Both the Spanish and the Quechua names are derived from the Quechua words “qhucha” (meaning "small lake") and “pampa” (meaning "open plain") ( Ajacopa 2007). We use the name " quchapampa " as a compound, proper noun (Latinized, fem.), in apposition to the genus name.

Diagnosis.

Ungla quchapampa is one of several species that has a prominent dark mark in the center of the frons, but it is the only one that has a dark flagellum and also lacks a prominent protuberance on the frons. The size and shape of the marks on the vertex are also distinctive: they are very large, dark brown, contiguous with each other, and extend anteriorly to the mesal margins of the antennal base. In more than half of the specimens we examined, the first intramedian cell (im1) of the forewing was quadrate (unlike the commonly triangular im1 cells of other Ungla species).

Ungla quchapampa closely resembles U. diazi in many features - notably size, wing features, and male abdominal characters (external and internal). In addition, the two species are known to occur sympatrically; however, U. quchapampa can be differentiated by its prominent frontal marking, sometimes quadrate im1 cell in the forewing, and the absence of markings on the ventral surface of the scape.

Description.

Head cream-colored with dark brown to black markings; vertex with inverted U-shaped marking large, robust, with lateral arms meeting anteriorly, extending forward into dorsal fossa, between antennae; mesal side, posterior margin of dorsal fossa marked with dark brown; posterior parts of vertex unmarked; frons with large, prominent mesal mark; clypeus with lateral margin black; tentorial pits surrounded by dark brown; gena with black stripe from eye, contiguous with black mark on clypeal margin. Antenna: scape cream-colored, dorsum with large, brown longitudinal mark; pedicel, flagellum dark brown to black basally, becoming lighter distally, with cream-colored intersegmental membrane; maxillary palp with basal two segments pale, three distal segments black, articulations pale; labial palp with basal segment pale, distal two segments marked with black.

Thorax with distinct longitudinal cream-colored stripe mesally; prothorax with pair of broad, dark reddish brown stripes laterally; transverse furrow in posterior region of segment, extending to lateral margins; setae mostly elongate, pale. Measurements: head width: 1.3-1.5 mm; ratio head width: eye width: 2.1-2.6: 1; prothorax width: 1.1-1.2 mm, length: 0.6-0.7 mm.

Forewing, hindwing somewhat broad, apex round; membrane clear, hyaline, without fumose area; stigma slightly opaque to light brown; longitudinal veins green, marked with brown at intersections with transverse veins; transverse veins mostly brown or marked with brown; forewing with veins robust, not crassate; Rs straight; first intramedian cell ovate or quadrate; basal inner gradate either meeting Psm or not; gradate veins, subcostal crossveins below stigma dark brown, costal crossveins, R-Rs crossveins, intracubital crossveins entirely brown; hindwing with costal crossveins, subcostal crossveins, gradate veins brown. Forewing 13.0-14.3 mm long, 4.6-5.1 mm wide [ratio L: W = 2.7-2.8: 1]; height of tallest costal cell 1.0-1.1 mm (cell number 5-6); width of first intramedian cell 1.2 mm (whether quadrate or ovate); 11-12 radial cells (closed cells between R and Rs); third gradate cell 1.5-1.6 mm long, 0.5 mm wide (ratio, L: W = 3.0-3.4: 1); fourth gradate cell 1.5-1.6 mm long, 0.4-0.5 mm wide (ratio, L: W = 3.1-3.8: 1); 3-4 Banksian cells (b cells, sometimes not distinct), 4 b’ cells; 5-7 inner gradates (basal one sometimes not distinct), 7 outer gradates. Hindwing 11.6-13.0 mm long, 3.9-4.3 mm wide (ratio, L: W = 2.9-3.0: 1), 10-11 radial cells, 3 Banksian (b) cells, 4 b’ cells, 5-6 inner gradates, 6-7 outer gradates.

Male. Abdomen with small spiracles (e.g., A7: spiracle diameter ~0.04 × length of sternite); subanal plate substantial, with ~20 setae in a well defined group; T9+ect relatively long (~0.6 × length of T7), with dorsal invagination deep (~0.7 × dorsal length of T9+ect), margins of invagination almost straight, base acute rounded; dorsal margin of T9+ect short, straight, angled downward to round distal tips; posterior margin of ectoproct rounded, protruding somewhat, posteroventral corner rounded but with small distal projection of apodeme, small knob; ventral margin bent, with moderately heavy apodeme except anteriorly; callus cerci large, ovate, margin sclerotized throughout; sclerotization probably contiguous with that on ventral margin of ectoproct. S8+9 fused, with line of fusion not perceptible; basal three-fourths of dorsal margin with heavy apodeme; dorsum tapering slightly anteriorly, forming concave depression at about 3/4th distance to tip; distal one-fourth forming concave terminus (lateral view), extending distally beyond T9+ect; setae slender, mostly long, simple, probably without flanges (We could not find any flanged setae; however, a number of the distolateral setae were missing). Gonarcus arcuate, V-shaped (frontal view), with bridge robust, bent mesally, arms elongate, extending ventrodistally from gonarcal bridge, rounded distally, mesal section with digitiform process extending posteromesally; mediuncus flat, closely attached to gonarcal bridge, not particularly long, extending distally or downward, ending in blunt, bent knob; bilobed gonosaccus, with lobes well separated, each lobe cylindrical, with rigid patch of large gonosetae facing mesally when uneverted, outwardly when everted; gonosetae arising from enlarged setal bases; hypandrium internum not found.

Known distribution.

BOLIVIA (central): Department of Cochabamba (Provinces of Carrasco and Chapare).

Specimens examined

(in addition to holotype). Same data as holotype (1F, paratype, MCZ); Bolivia, Cochabamba, Carrasco, Siberia 1650 m, xxii.1962-i.1963, i.1964, F. H. Walz (5F, all paratypes, CAS). Four of the specimens in the CAS have attached notes handwritten by P. A. Adams.