Vladimiretskia nathani Fedotova et Perkovsky, 2024

Fedotova, Zoya A., Vasilenko, Dmitry V. & Perkovsky, Evgeny E., 2024, Vladimiretskia nathani gen. et sp. nov. (Diptera, Cecidomyiidae: Porricondylinae, Asynaptini, Vladimiretskiana subtr. nov.) from Eocene Rovno amber with description of their eggs, Ecologica Montenegrina 72, pp. 146-157 : 149-157

Vladimiretskia nathani Fedotova et Perkovsky , sp. nov.

https://zoobank.org/ urn:lsid:zoobank.org:act:5BC8687E-0758-4D08-9FFB-EDBF10F10BA8

( Figs 1-3 )

Material: SIZK W-73, holotype, Vladimirets, Rovno amber, late Eocene. Well preserved female inclusion with tarsi separated and partially lost. Seven eggs in two groups are located near the body. Syninclusion : SIZK W-74, Сollembola, Entomobryomorpha .

Description. Female (figs 1 b, с, e–h; 2 a–g; 3 a–j). Body dark-brown, ovipositor 2.5 times longer than antennae, 1.1 times as long as wing. Antennae 2.3 times shorter than wings, 2.2 times longer than lateral side of head. Head with large eyes, width 1.5 times as long as head height. Pedicel slightly broad, almost as long as scapus, 1.9 times shorter than 1 st flagellomere. 2 nd flagellomere almost as long as 1 st. 1 st- 3 rd flagellomeres without neck; 5 th flagellomere almost 2.0 times as long as wide, 1.1 times as long as 1 st flagellomere, basal node slightly narrowed basally, enlarged distally, 4.6 times as long as neck. 16 th flagellomere narrowed apically, slightly longer 15 th, 1.1 times shorter than 5 th. The dark palpiger almost round, palpi very narrow, 1 st –3rd palpomeres parallel laterally. 3 rd palpomere strongly prolonged, 2.4 times as long as 2 nd. Wing 2.6 times as long as wide. R 1 almost 2.0 times shorter than C. Fore femur almost as long as fore tibia. Ratio of fore tarsi 1:9.1:4.6:3.0:1.5. Middle femur 1.1 times length of fore femur, 1.1 times length of middle tibia. Ratio of middle tarsi 1:9.3:4.8:3.1:1.6. Middle tarsi 1.6 times length of fore tarsi. Hind femur 1.3 times length of fore femur. Ratio of hind tarsi 1:12.1:6.0:6.9: 5 th is not preserved. Hind tarsi 1.3 times length of fore tarsi. Apical lamelles of ovipositor 1.1 times as long as 9 th abdomen segment.

Eggs (figs 1 a, d, f). Seven eggs in two groups were found in the amber sample next to the female (3 + 4). They are oval, slightly expanded, very slightly curved in the middle, and the apices are widely rounded. All are shiny and appear black seen from one side of the amber piece, but are transparent seen from the other side. Their length and width vary slightly: length 0.423 –0.494 mm, width 0.082 – 0.105 mm. Their lengths are 4.2–5.3 times greater than widths.

Measurements (mm): body length, 3.18; antennal length, 1.26; head height, 0.32; width of lateral side of head, 0.47; length of head front, 0.57; width of frontal side of head, 0.47; palpus length, 0.20; thorax length, 0.72; thorax width, 0.77; wing length, 3.01; wing width, 1.06; R 1, 1.38; halter length, 0.45; halter knob, 0.17 mm; length of fore leg, 4.67: coxa, 0.29; trochanter, 0.12; femur, 0.96; tibia, 0.93; 1 st tarsomere, 0.12; 2 nd, 1.12; 3 rd 0.57; 4 th 0.37; 5 th 0.19; fore tarsi, 2.37; length of middle leg, 3.74 without coxa and trochanter: femur, 1.05; tibia, 0.93; 1 st tarsomere, 0.10; 2 nd, 0.63; 3 rd 0.50; 4 th 0.33; 5 th 0.20; middle tarsi, 1.76; length of hind: femur, 0.74; tibia, 0.51; 1 st tarsomere, 0.123; 2 nd, 0.70; 3 rd 0.34; 4 th 0.39; 5 th lost; abdomen length, 2.06; length of 1-9 segments, 1.59; length of ovipositor (3 apical plates), 0.18.

Eggs (length: width, mm): 1). 0.494: 0.105; 2). 0.442: 0.084; 3). 0.442: 0.105; 4). 0.423: 0.082; 5). 0.435: 0.094; 6). 0.435: 0.094.

Etymology. The species is named for Nathan Perkovsky, the late son of the third author.

Acknowledgements

Authors are thankful to Alexandr P. Rasnitsyn (Paleontological Institute, Moscow, Russia) for the discussion of the first version of the manuscript, S. Bruce Archibald (University of British Columbia) for editing the English and reviewers for their useful comments.

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Figures

Figure 1. Vladimiretskia nathani Fedotova et Perkovsky sp. nov. (holotype, female, Rovno amber, SIZK W-73, photomicrographs): a, d, f – eggs; b – general view, laterally and eggs; c – palpi; e – part of abdomen; g – ovipositor; h – 6 and 7 flagellomeres; 1–7 -аrrangement of eggs by numbers. Scales = 1.3 mm (b); 0.1 mm (a, dh); 0.2 mm (c).

Figure 2. Vladimiretskia nathani Fedotova et Perkovsky sp. nov. (holotype, female, Rovno amber, SIZK W-73, photomicrographs): a – head, palpi, antennae proximally, thorax, halter, fore coxa and trochanter; b – head, palpi and antennae frontally; c – fore tibia distally and 1st tarsomere with lateral spiniform projection; d – left wing and halters; e – 5th mesotarsomere with tarsal claw dorsally; f – 5th fore tarsomere with claw laterally; g – 6-16th flagellomeres. Scales =1.0 mm (d); 0.5 mm (b); 0.2 mm (a, c, e, f, g).

Figure 3. Vladimiretskia nathani Fedotova et Perkovsky sp. nov. (holotype, female, Rovno amber, SIZK W-73, drawings): a – wing; b – 13–16th flagellomeres; c – 6–7th flagellomeres; d – palpi; e – fore tarsus; f – 1st fore tarsomere; g –5th mesotarsomere dorsally; h – 5th fore tarsomere laterally; i – apex part of head with eye, scape, pedicel and 1-3rd flagellomeres; j – IX abdominal segment and cerci (ovipositor). Scales = 0.1 mm (d, f); 0.2 mm (e); 0.25 mm (a).