Wakarumbia pseudofasciata, Dvorak & Bocak, 2009

Dvorak, M. & Bocak, L., 2009, Ten new species of Wakarumbia Bocak, 1999 from Sulawesi (Coleoptera: Lycidae), with a key to males of the genus, Zootaxa 2282 (1), pp. 51-61 : 59-61

Wakarumbia pseudofasciata sp. nov.

(Figs 19–20)

Type material. Holotype. Male, W. Sulawesi, 8 km W of Mamasa (Nepe), 950 m, 119.20.32E 2.56.13S, 29.– 31. June 2001, Bolm lgt. ( LMBC) . Paratypes. 2 males, 2 females, ditto ; 2 males and 1 female, W. Sulawesi, 25 km E of Mamasa (Kalama), 1100 m, 119.28.39E 3.02.10S, 1.–3. July 2001, Bolm lgt. ( LMBC) .

Differential diagnosis. W. pseudofasciata sp. nov. is similar in the body form and color pattern to syntopically occurring W. kalamensis and W. fasciata . Close examination of male genitalia is necessary for a reliable identification of these species. W. pseudofasciata is distinguishable by a very short robust tubular part of the phallus (Figs 19–20).

Description. Body medium-sized, thorax and head dark brown to black, elytra black with transverse yellow patch in humeral third of their length. Head small, partly hidden by pronotum, eyes large, hemispherically prominent, frontal interocular distance 1.10 times maximum eye diameter. Antennae compressed, antennomere 3 2.2 times longer than width. Pronotum flat, 1.32 times wider than long, with well developed carinae, disc uniformly black, narrow area along frontal and lateral margins densely punctured, whole disc covered with short, fine, dense pubescence. Elytra parallel-sided, very slender, 4.16 times longer than width at humeri, reticulate cells regular, mostly quadrate. Phallus stout in basal part, slender in middle, apical part slightly rotated (Figs 19–20).

Measurements. BL 7.43 mm, HW 1.50 mm, PL 0.97 mm, PW 1.28 mm, EL 6.25 mm, Ediam 0.48 mm, Edist 0.53 mm.

Distribution. Sulawesi, the Mamasa valley.

Etymology. The specific epithet points toward similarity of W. fasciata and W. pseudofasciata .

Key to males of the genus Wakarumbia *

*Only males can be identified by the key. The females of many species cannot be identified by the external morphology and coloration.

1. Elytra dark brown to black, at most with slightly lighter costae .................................................................................. 2

- Elytra otherwise colored, pale yellow to light brown, black with yellow humeri, or black with transverse band in middle part of elytra ................................................................................................................................................... 18

2. Head, thorax and most of femora light brown, pronotum brown, phallus symmetrical, Buton Isl. .............................. ................................................................................................................................................. W. gracilis Bocak, 1999

- Body dark brown to black, only trochanters and pronotal margins light brown or eventually elytral costae a slightly lighter than bottom of elytral cells, Sulawesi ............................................................................................................... 3

3. Eye diameter at least 1.10 times longer than interocular distance................................................................................ 4

- Eye diameter smaller than interocular distance or their diameter equals interocular distance .................................... 8

4. Eye diameter 1.10–1.27 times longer than interocular distance, apical part of phallus symmetrical .......................... 5

- Eye diameter at least 1.50 times longer than interocular distance, phallus slightly rotated apically .......................... 7

5. Pronotum dark brown, wide frontal margin of pronotum light brown, lateral margins and basal parts of costae brown, eye diameter 1.17 times interocular distance, phallus robust at base (Figs 9–10), body pubescence brown .... ...................................................................................................................................................... W. kundratai sp. nov.

- Pronotum completely black .......................................................................................................................................... 6

6. Basal tubular part of phallus occupies more than quarter of phallic length, maximum eye diameter 1.4 interocular distance, pubescence black ....................................................................................................... W. oculata Bocak, 2000

- Basal tubular part of phallus occupies less than one fifth of phallic length, maximum eye diameter 1.2 interocular distance, pronotum and elytra covered with dense, long gray pubescence ................................ W. grisea Bocak, 2001

7. Basal tubular part of phallus occupies about a third of phallic length............................... W. kalamensis Bocak, 2001

- Basal tubular part of phallus occupies about one tenth of phallic length (Figs 11–12).................. W. linearis sp. nov.

8. Phallus with two separate spines in internal sac (Figs 5–6) .............................................................. W. hanae sp. nov.

- Phallus with one complex spine in internal sac (Figs 7–8) .......................................................................................... 9

9. Apical part of phallus considerably rotated (Figs 7–8) .............................................................................................. 10

- Apical part of phallus symmetrical or rotated by less than 90 degrees (Figs 3–4) .................................................... 12

10. Tubular part of the phallus represents more than 30 percent of phallus length, interocular distance 1.15 longer than eye diameter, body large, around 7 mm, very narrow margin of pronotum brown ................. W. grandis Bocak, 2000

- Tubular part of the phallus represents less than 20 percent of phallus length ............................................................ 11

11. Tubular part of phallus about twice longer than width of basal part, completely black colored species ...................... ...................................................................................................................................................... W. petri Bocak, 2001

- Tubular part of the phallus about as long as width of basal part of phallus, dark brown colored species (Figs 7–8) ... ......................................................................................................................................................... W. hirsuta sp. nov.

12. Elytral costae lighter than bottom of elytral cells, phallus widest in basal third ...................... W. similis Bocak, 2000

- Elytra costae of same color as bottom of elytral cells ............................................................................................... 13

13. Tubular part of phallus very long, it forms about 40 percent of phallus length (Figs 3–4) ............ W. decepta sp. nov.

- Tubular part of phallus forms at most a quarter of phallic length ............................................................................. 14

14. Length of tubular part of phallus about twice longer than width of basal part.......................................................... 15

- Length of tubular part of phallus equals width of basal part or shorter...................................................................... 16

15. Phallus almost symmetrical, at most apical part slightly rotated (less than 30 degrees), interocular distance 1.25 times longer than maximum eye diameter ............................................................................. W. montana Bocak, 2001

- Phallus asymmetrical, apical part of phallus rotated by about 60 degrees, interocular distance 1.36 times maximum eye diameter ....................................................................................................................... W. orobuensis Bocak, 2001

16. Eye diameter approximately equals interocular distance ............................................................ equalis Bocak, 2001

- Eye diameter about 0.8 interocular distance ............................................................................................................... 17

17. Pronotum with light gray, long, dense pubescence, tubular part of phallus shorter than width of basal part (Figs 13– 14) .................................................................................................................................................. W. obscura sp. nov.

- Pronotum with dark pubescence, tubular part of phallus about as long as width of basal part .. W. nigra Bocak, 2000

18. Elytra black with yellow transverse band in middle, sometimes yellow part can be diminished and forms more or less independent bright patches on each elytron ......................................................................................................... 19

- Elytra uniformly light brown, with light humeri or uniformly brightly yellow ......................................................... 22

19. Phallus robust basally, tubular part very short (Figs 17–18) .................................................. W. pendolensis sp. nov.

- Phallus slender on whole length, tubular part represents at least one fifth of phallic length (Figs 17–18) ............... 20

20. Tubular part of phallus long, representing two fifths of phallic length ............................. W. mamasensis Bocak, 2001

- Tubular part short, occupying at most one third of phallic length ............................................................................. 21

21. Base of phallus robust (Figs 19–20) ................................................................................... W. pseudofasciata sp. nov.

- Base of phallus slender ............................................................................................................ W. fasciata Bocak, 2001

22. Whole elytra brown, gray, light brown to pale yellow or at most apical one tenth of elytra slightly infuscate ........ 23

- Humeral part of elytra lighter than apical one, infuscate part always larger than light one ......................................

23. Basal tubular part of phallus, slender, parallel-sided, tubular part forms about 40 percent of phallic length ............... ................................................................................................................................................ W. brendelli Bocak, 2000

- Basal tubular part of phallus short, tubular part forms 5–10 percent of phallic length .............................................. 24

24. Cranium yellow, elytra yellow with black apex, male genitalia with wide apical part (Figs 1–2) .... W. aurea sp. nov.

- Cranium black or brown, always darker than adjacent part of pronotum, elytra uniformly yellow or at most slightly darker brown at apex.................................................................................................................................................. 25

25. Phallus apparently asymmetrical, apical part of phallus rotated at least by 90º .................. W. pallescens Bocak, 2000

- Apical part of phallus almost symmetrical (Figs 15–16) ........................................................................................... 26

26. Tubular part of phallus much shorter than width of phallus in basal part ................................................................. 27

- Tubular part of phallus only slightly shorter than width of phallus in basal part ............ W. amporivensis Bocak, 2001

27. Maximum eye diameter 1.2 frontal interocular distance ................................................... W. brunescens Bocak, 2000

- Maximum eye diameter 1.4 frontal interocular distance ............................................................ W. obstinata sp. nov.. At least one quarter of elytra yellow .......................................................................................................................... 29

- Only very short transverse part of humeri lighter than the rest of elytra ................................................................... 30

29. Elytral costae slightly lighter than bottom of cells in humeral third of elytra, male eye diameter 1.67 times interocular distance, phallus without any constriction in apical part ......................................... W. flavohumeralis Bocak, 2000

- Coloration of elytral costae and bottom of elytral cells similar in whole elytra, male eye diameter 1.35 times interocular distance, phallus with considerably constricted in subapical part ...................................... W. angustior Pic, 1922

30. Disc of pronotum infuscate, elytral cells apparently transverse, eyes in male as large as interocular distance ............ ............................................................................................................................................ W. celebensis Kleine, 1927

- Disc of pronotum yellow, without any patches, elytral cells slightly transverse ......................... W. lateniger Pic, 1922