Zhiganka longialata, Lukashevich, 2019

Lukashevich, Elena D., 2019, Review of the fossil record of Bittacomorphinae (Diptera: Ptychopteridae), Zootaxa 4661 (3), pp. 566-578 : 568-573

Zhiganka longialata sp.nov.

urn:lsid:zoobank.org:act:BF7F96FE-AACA-49DA-AC44-0E8096F32EA4

Holotype. PIN, No. 5340/953, part and counterpart, well-preserved impression of a male with detached terminalia.

Type Horizon and Locality. Khasurty; Western Transbaikalia, Buryatia, Zakamensk District, middle reaches of the River Khasurty 10 km south of the village of Tsakir (50°21’N 103°37’E); lower part of Gusinoe Ozero Group, Lower Cretaceous.

Etymology. The species epithet refers to relatively long wings of the species (from Latin longus —long and alata —winged).

Description. Antenna subequal to wing in males and shorter than wing in females ( Fig. 1B ). Wing 3.0–3.4x as long as wide, transparent, with single pale pigmented spot, pterostigma ( Fig. 4A ). Sc ends well beyond midwing and beyond CuA termination. Rs originates at about one-quarter of wing length. R 2 well distal to R 4+5 furcation, distal section of R 1 subequal to R 2, R 2+3 2.4x as long as R 3. R 4+5 fork 1.3x as long as R 4+5 stem, R 4+5 stem subequal to Rs stem ( Figs 2B , 3C , 5A ). Hind tibia longest of leg segments. Epandrium with broad and apically rounded setose surstyli ( Figs 2E, F , 3 F–H, 4D, E).

Comparison. In its wing size and position of R 2 and r-m the new species is closer to Z. comitans ( Fig. 5A, B ), differing in a shorter Rs (75% of the wing length instead of 81% in the type species, according to specified data) and shorter R 2+3 (2.4x as long as R 3 instead of 3x).

Measurements (mm). Male (n=6). Body length (from anterior region of head to abdomen tip) 9.4–11.6, lateral thorax length 1.4–1.7, abdomen length 7.4–9.2. Wing length 6.8–8.4, width 2.3–2.4. Antenna length (n=3) 5.2–7.2. Leg segment length in Table 1. Female (n=5). Body length 11.0–12.6, abdomen length 9.0–9.8. Wing length 8.8– 9.6, width 3.0. Antenna length (n=1) 4.0. Hind leg: femur 4.2, tibia 5.6, tarsus 4.2 (2.6:0.8:0.5:0.2:0.1)

Leg segment length in holotype PIN 5340 /953, male (in mm) Additional material. Paratypes PIN, No. 5026/1594±,1598±, 1599± and 5340/764±, 807±, complete males ; PIN, No. 5026/1593±, 1595±, 1597± and 5340/951±, 952±, 1643± partly preserved females; PIN, No. 5026/1600 and 5340/1494± partly preserved adults, sex unknown; PIN, No. 5026/1596± isolated wing.

Figures

FIGURE 1. Zhiganka longialata sp. nov.: A–E. Male PIN 5340/953, holotype:A. Head and thorax. B. Habitus. C. Two spurs on hind tibia. D, E. Two spurs on left and right midtibia. F. Spur on fore tibia, PIN 5340/1494. G. Habitus of female PIN 5340/951, paratype. H. Terminalia of female PIN 5026/1595. Abbreviations: cl, clypeus; lbl, labellum; lbr, labrum; mp, maxillary palp; s, scutal suture; sl, scutellum.

FIGURE 2. Zhiganka longialata sp. nov., male PIN 5026/1599, paratype: A. Anterior part of body. B. Wing. C. Proximal flagellomeres of antenna, inset—distal flagellomeres, arrows mark setae. D. Head. E, F. Terminalia, part and counterpart. G, Distal part of wing. Abbreviations: apn, antepronotum; cl, clypeus; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ss, surstylus.

FIGURE 3. Zhiganka longialata sp. nov.: A–C. Paratype PIN 5026/1599: A. Anterior part of body. B. Distal and proximal flagellomeres of antenna. C. Wing. D. Abdomen of female PIN 5026/1597, dorsal view. E. Abdomen of male PIN 5340/807, lateral view (flipped horizontally). F–H. Genitalia of male. F. Paratype PIN 5026/1599. G. Paratype PIN 5026/1594. H. Paratype PIN 5340/807. Scale bars equal 2 mm (A, C, D, E), 200 μm (B, F, G, H). Abbreviations: apn, antepronotum; b, transepimeral suture; cl, clypeus; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ps, prescutal suture; s, scutal suture; ss, surstylus.

FIGURE 4. Zhiganka longialata sp. nov.: A, B. Anterior and basal part of wing, PIN 5340/1494, part and counterpart. C. Part of wing, female PIN 5340/951. D. Genitalia of male PIN 5026/1594. E. Genitalia of male PIN 5340/953. F. Thorax and head, female PIN 5340/952. Abbreviations: aem, anepimerum or pteropleurite; apn, antepronotum; b, transepimeral suture; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lc, laterocervicale; mt, mediotergum, mpl, meropleurum; ps, prescutal suture; pt, pleurotergum; s, scutal suture; sl, scutellum; ss, surstylus.

FIGURE 5. SEM images:A. Zhiganka longialata sp. nov., holotype PIN 5340/953, wing; Khasurty, K 1. B. Zhiganka comitans, holotype PIN 4502/13, Zhigansk, K 1. C. Crenoptychoptera antica, PIN 1255/2123, wing; Kubekovo, J 2. Arrows mark spurious vein. Scale bars equal 2 mm.