Cytisus frivaldszkyanus Degen in Oesterr. Bot. Z. 43: 422 (1893)

Sennikov, Alexander N. & Tikhomirov, Valery N., 2024, Atlas Florae Europaeae notes, 35. Further critical notes on Cytisus sect. Tubocytisus (Fabaceae) in Europe, PhytoKeys 238, pp. 199-230 : 199

7. Cytisus frivaldszkyanus Degen in Oesterr. Bot. Z. 43: 422 (1893)

- Chamaecytisus frivaldszkyanus (Degen) Kuzmanov in Jordanov, Fl. Narodna Republ. Bulg. 6: 110 (1976); Kuzmanov in Taxon 24: 504 (1975), comb. inval. (Art. 41.1).

= Cytisus microphyllus Boiss., Diagn. Pl. Orient., ser. 2, 2: 5 (1856), non Link (1825), nom. illeg. (Art. 53.1), syn. nov. - Cytisus austriacus var. microphyllus Boiss., Fl. Orient. 2: 53 (1872) - Cytisus austriacus subsp. microphyllus (Boiss.) Cristof. in Webbia 45(2): 210 (1991). Type. Greece. "In monte Pelione", P. Aucher-Éloy 1109 (holotype G; isotypes BM 000750890, K 000829488).

= Cytisus rhodopeus J.Wagner ex Bornm. in Bot. Jahrb. Syst. 59(5): 465 (1925) - Chamaecytisus absinthioides subsp. rhodopeus (J.Wagner ex Bornm.) Kuzmanov in Taxon 21: 336 (1972), comb. inval. (Art. 41.1) - Chamaecytisus absinthioides var. rhodopeus (J.Wagner ex Bornm.) Micevski, Fl. Macedon. 1(5): 1140 (2001), comb. inval. (Art. 41.1). Type. Bulgaria. "In graminosis decliv. m. Osogovska Planina", 08.1887, J. Velenovský (PRC 456104, lectotype designated here). Fig. 10 .

- Cytisus lasiosemius auct.: Cristofolini (1991).

- Chamaecytisus supinus subsp. lasiosemius auct.: Niketić (2021).

Type.

Bulgaria. "In declivibus dumetosis montis Rhodopes centralis pr. Stanimak (inter Hvojna et Bačkova)”, 06.1892, J. Wagner 39 (syntypes JE, PRC); "In declivibus dumetosis prope Slivno (Balkan)", 07.1893, J. Wagner (syntypes JE, PRC); "In dumetosis montis “Čatal " prope Slivno", 21.07.1893, J. Wagner 31 (syntype PRC); "Bela Cerkva", Skorpil (syntype not traced) .

Distribution.

Europe: Balkan Peninsula (Bulgaria, Greece, North Macedonia, Serbia) ( Kuzmanov 1976; Micevski 2001; Assyov and Petrova 2012; Barina et al. 2018; Niketić 2021) (Fig. 11 ). This species was reported from Albania ( Barina et al. 2018), but the background of this report has not been examined by us.

Notes on taxonomy.

This species with subpatent to patent pubescence was accepted by Cristofolini (1991), but under a wrong name, C. lasiosemius , probably because of the unavailability of the type collection of the latter species name.

Notes on nomenclature.

Degen (1893) described Cytisus frivaldszkyanus from a few localities in present-day Bulgaria, citing four syntype gatherings. The examined material is fairly homogeneous, and the application of the species name is unambiguous. So far, we refrain from lectotypification because the main collection of Degen at BP has not been examined by us.

Cytisus rhodopeus was first mentioned in the synonymy of C. eriocarpus by Degen (1893) and validly published by Bornmüller (1925) without any descriptive matter, but with a reference to the description of C. absinthioides in Velenovský (1891). Five syntypes from Bulgaria were cited in the original description ( Velenovský 1891), which deviated much from the description of the true C. absinthioides provided by Janka (1872) by a longer calyx (13-15 mm long vs. 7-8 mm long in C. absinthioides ) with patent (vs. appressed) hairs.

Through the kindness of P. Mráz, we traced a specimen in the collection of J. Velenovský at PRC, which exactly corresponds to the protologue by its diagnostic characters and taxonomic references on its label (to C. absinthioides Janka and " C. eriocarpus Boiss. var.", as Velenovský (1891) also noted a relationship with the latter species). This specimen fully reflects the taxonomic concept of Velenovský (1891) and is designated as a lectotype of C. rhodopeus here.

Cristofolini (1991) accepted C. austriacus subsp. microphyllus "(Boiss.) Boiss." as the correct name for a small-leaved segregate of C. austriacus , citing C. pindicola (Degen) Halácsy in its synonymy. The type collection of C. microphyllus Boiss. is quite dissimilar from C. pindicola and belongs to C. frivaldszkyanus because of its strong suberect stems, partly obovate (vs. lanceolate) leaflets and pods with nearly patent (vs. appressed) hairs.

Figures

Figure 10. Lectotype of Cytisus rhodopeus J. Wagner ex Bornm.

Figure 11. Distribution of Cytisus frivaldszkyanus Degen.