Exocelina jaseminae (Balke, 1998)

Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo & Balke, Michael, 2019, Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 878, pp. 73-143 : 73

11. Exocelina jaseminae (Balke, 1998) Figs 27 , 31

Copelatus (Papuadytes) jaseminae Balke, 1998: 331; Nilsson 2001: 77 (catalogue).

Papuadytes jaseminae (Balke, 1998): Nilsson and Fery 2006: 56 (comb. nov.).

Exocelina jaseminae (Balke, 1998): Nilsson 2007: 33 (comb. nov.).

Exocelina jaseminae MB1382: Toussaint et al. 2014: supplementary figs 1-4, tab. 2; Toussaint et al. 2015: supplementary figs S1, S2, tab. S3, and information S5, S6.

Type locality.

Papua New Guinea: Morobe Province, Herzog Range, Wagau (Vagau), ca. 06°48'S, 146°48'E, ca. 1300 m a.s.l.

Type material studied.

Holotype: male "Stn. No. 150", "NEW GUINEA: Morobe Dist., Herzog Mts., Vagau, C.4,000ft. 4 –17.i.1965”, "M. E. Bacchus. B. M. 1965-120", “HOLOTYPUS” [red], " Copelatus jaseminae sp. nov. Balke des. 1997" [red] (BMNH). Paratypes: 2 males, 3 females with the same label as the holotype and additionally with a red label "Paratypus Copelatus jaseminae sp.n. Balke des. 1997", one of the males with an additional label "measured J. Parkin 85" (BMNH, NHMW). 1 male with the same label as the holotype, the red paratype label missing (BMNH).

Additional material.

Morobe: 1 male "Stn. No. 149A", "NEW GUINEA: Morobe Dist., Herzog Mts., Vagau, C.4,000ft. 4 –17.i.1965”, "M. E. Bacchus. B. M. 1965-120", "Paratypus Copelatus monae sp.n. Balke des. 1997" [red], " Exocelina jaseminae (Balke) det. H.Shaverdo 2014" (BMNH). 2 males "Papua New Guinea: Morobe, Wagau, Herzog Mts., 1150m, 19.xi.2006, 06.51.067S 146.48.068E, Balke & Kinibel (PNG 102)", one male with an additional green label "DNA M.Balke 1382" (ZSM). 1 male, 2 females "Papua New Guinea: Morobe, Wagau, Herzog Mts., 1150m, 19.xi.2006, 06.51.067S 146.48.068E, Balke & Kinibel (PNG 103)" (ZSM). 2 males "Papua New Guinea: Gulf [sic!], Menyamya, Mt Inji 1700m, 14.xi.2006 nr 07.14.813S 146.01.330E Balke & Kinibel (PNG 96)" (ZSM). 9 males "PAPUA N. G.: Morobe Prov. Aseki, Oiwa, 1600-1700 m, 11.-12.3.1998 leg. A. Riedel" (NHMW). 6 males, 2 females "Papua New Guinea: Gulf [Morobe], Marawaka, Andakombe towards Morobe, 1100m, 12.xi.200, 07.09.766S, 145.46.333E, Balke & Kinibel (PNG 92)" (NHMW, ZSM). Eastern Highlands: 38 males, 38 females "Papua New Guinea: Eastern Highlands, Marawaka, Ande, 1700m, 8.xi.2005, 07.01.697S 145.49.807E, Balke & Kinibel (PNG 86)", one male with an additional green label "DNA M.Balke 1365" (NHMW, ZSM). 49 males, 40 females "Papua New Guinea: Eastern Highlands, Marawaka, Ande, 1700-1800m, 9.xi.2006, 07.01.697S 145.49.807E, Balke & Kinibel (PNG 87)" (NHMW, ZSM).

Diagnosis.

For complete description, see Balke (1998: 331). Beetle medium-sized: TL-H 3.55-4.1 mm; oblong-oval; brown to dark brown, with reddish to reddish brown pronotal sides or pronotum and often head; shiny, with very fine, on elytra often almost invisible punctation and weakly impressed microreticulation; pronotum with lateral bead ( Fig. 27 ); male protarsomere 4 with anterolateral seta thin, evenly curved, smaller than or equal to more laterally situated large seta; male protarsomere 5 ventrally with anterior band of more than 60 and posterior row of 13-15 relatively long setae ( Fig. 31D ); median lobe in lateral view with apical lobes distinctly developed and rounded, “nose” small but distinct; paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin distinctly divided to dense and strong subdistal setae and sparser proximal ones, setae in middle short and fine ( Fig. 31 A–C ).

Affinities.

In the area of its distribution, E. jaseminae co-occurs with numerous species of the E. ekari , E. ullrichi , E. broschii , and E. danae groups. From them, this species can be distinguished by its size, dorsal colouration, surface sculpture, simple male antennae, presence of pronotal bead, and mainly by the shape of its median lobe. In its external appearance, E. jaseminae is especially similar to E. monae (Balke, 1998), from which can be distinguished by the shape of its median lobe. For the affinities within the group, see the “Key”.

Distribution.

Papua New Guinea: Eastern Highlands and Morobe Provinces ( Fig. 34 ).

Figures

Figure 34. Map of the eastern part of New Guinea showing distribution of the species of the E. jaseminae group.

Figures 26 - 29. Habitus and colouration 26 Exocelina aseki sp. nov. 27 E. jaseminae (Balke, 1998) 28 E. kailaki sp. nov. 29 E. pseudojaseminae sp. nov.

Figures 30, 31. 30 Exocelina aseki sp. nov. 31 E. jaseminae (Balke, 1998) A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view.