Ochyrocera varys, Brescovit, Antonio D., Cizauskas, Igor & Mota, Leandro P., 2018

Brescovit, Antonio D., Cizauskas, Igor & Mota, Leandro P., 2018, Seven new species of the spider genus Ochyrocera from caves in Floresta Nacional de Carajas, PA, Brazil (Araneae, Ochyroceratidae), ZooKeys 726, pp. 87-130 : 88-97

Ochyrocera varys sp. n. Figures 1, 2, 3, 19A, 21A, D

Types.

Holotype male from Cave N5 S– 0021 (6°5'15"S, 50°7'34"W), Serra Norte, Floresta Nacional de Carajás, Parauapebas, Pará, Brazil, 25/ VIII– 03/IX/2009, R. Andrade & I. Cizauskas (IBSP 177662). Paratypes: female from Cave N4 E– 0033 (6°2'25"S, 50°9'36"W), Serra Norte, Floresta Nacional de Carajás, Parauapebas, Pará, Brazil, 15-22/IX/2009, R. Andrade & I. Cizauskas (IBSP 176843); male and female from N5 S– 0085 (6°5'12"S, 50°7'35"W), Serra Norte, Floresta Nacional de Carajás, Parauapebas, Pará, Brazil, 25/ VIII– 03/IX/2009, R. Andrade & I. Cizauskas (MPEG 34434, ex IBSP 177644); 2 males, 2 females, N5 S– 0085 (6°5'12"S, 50°7'35"W), Serra Norte, Floresta Nacional de Carajás, Parauapebas, Pará, Brazil, 14/ III– 04/IV/2010, R. Andrade & I. Cizauskas (MZSP 72854, ex IBSP 177735).

Other material examined.

BRAZIL. Pará: HYPOGEAN SAMPLES: Canaã dos Carajás, Floresta Nacional de Carajás, Serra Sul, Cave CAV_0017 (6°24'23"S; 50°22'9"W), 1♀, 22-31/V/2010 (IBSP 175566); Cave CAV_0034 (6°24'9"S, 50°22'56"W), 1♂, 22-31/V/2010 (IBSP 175316); Cave CAV_0018 (6°29'50"S, 51°9'31"W), 3♀, 08-15/III/2012 (IBSP 175979) all collected by R. Andrade & I. Cizauskas et al.; Cave S11A-03 (6°20'60"S, 50°27'2"W), 2♂7♀, 23/VIII−02/IX/2007 (IBSP 174480); Cave S11A-05 (6°21'7"S, 50°27'3"W), 3♂11♀, 23/VIII−02/IX/2007 (IBSP 174405); 1♂3♀, 06/IV/2017, R. Zampaulo & X. Prous (IBSP 194762); Cave S11A-07 (6°21'6"S, 50°26'36"W), 2♂3♀, 23/VIII−02/IX/2007 (IBSP 174410); Cave S11A-12 (6°19'53"S; 50°27'4"W), 1♂5♀, 23/VIII−02/IX/2007 (IBSP 174413); Cave S11A-20 (6°19'4"S, 50°26'23"W), 1♂5♀, 23/VIII−02/IX/2007 (IBSP 174416, IBSP 174417); Cave S11A-26 (6°18'26"S, 50°26'55"W), 3♂6♀, 23/VIII-02/IX/2007 (IBSP 174423, IBSP 174418); Cave S11A-36 (6°19'1"S, 50°27'17"W), 4♂4♀, 23/VIII−02/IX/2007 (IBSP 174429); Cave S11B-09 (6°21'19"S, 50°23'24"W), 1♂1♀, 23/VIII−02/IX/2007 (IBSP 174433); Cave S11B-11 (21'27"S; 50°23'22"W), 3♀, 23/VIII−02/IX/2007 (IBSP 174435); Cave S11B-13 (6°21'16"S, 50°26'51"W), 6♀, 23/VIII−02/IX/2007 (IBSP 174439); Cave S11B-14 (6°20'59"S, 50°24'10"W), 1♀, 23/VIII−02/IX/2007 (IBSP 174441); Cave S11B-23 (6°20'43"S, 50°24'35"W), 2♀, 23/VIII−02/IX/2007 (IBSP 174443); Cave S11B-24 (6°20'43"S, 50°°24'34"W), 3♀, 23/VIII−02/IX/2007 (IBSP 174446); Cave S11B-49 (6°21'31"S, 50°23'21"W), 2♂, 23/VIII−02/IX/2007 (IBSP 174448); Cave S11C-14 (6°24'10"S, 50°22'57"W), 3♂5♀, 23/VIII−02/IX/2007 (IBSP 174451); Cave S11C-27 (6°23'51"S, 50°23'21"W), 2♀, 23/VIII−02/IX/2007 (IBSP 174453) all collected by R. Andrade & I. Arnori et al.; Cave S11D-06 (6°24'4"S, 50°21'1"W), 1♂1♀, 19-22/II/2010 (IBSP 175534); Cave S11D-11 (6°23'51"S, 50°21'30"W), 1♀, 13-30/I/2010 (IBSP 175512); Cave S11D-43 (6°24'48"S, 50°19'17"W), 1♂2♀, 13-30/I/2010 (IBSP 175521); Cave S11D-64 (6°23'31"S, 50°18'48"W), 1♂1♀, 01-14/VII/2010 (IBSP 175653); Cave S11D-78 (6°23'32"S, 50°18'58"W), 1♀, 13-30/I/2010 (IBSP 175423) all collected by R. Andrade & I. Cizauskas et al.; Parauapebas, Cave CRIS-28 (6°27'31"S, 49°42'33"W), 2♀, 29/VII−06/VIII/2008, R. Andrade (IBSP 174624); Floresta Nacional de Carajás. Serra Norte, Cave GEM-1564, 10♀, 17-24/X/2008 (IBSP 174508); Cave GEM-1590, 5♀, 17-24/X/2008, R. Andrade (IBSP 174521); Cave N1_0002 (6°2'24"S, 50°16'12"W), 1♀, 28/IX-03/X/2007, R. Andrade & I. Arnori et al. (IBSP 174652); Cave N1_0004 (6°2'23"S, 50°16'12"W), 1♂8♀, 28/IX-03/X/2007, R. Andrade & I. Arnori et al. (IBSP 174655); 3♀, 16/VII-06/VIII/2014, Equipe Carste et al. (IBSP 188836, IBSP 188842); Cave N1_0008 (6°2'20"S, 50°16'13"W), 6♀, 28/IX−03/X/2007 (IBSP 174657, IBSP 174656); Cave N1_0014 (6°2'2"S, 50°16'20"W), 4♀, 28/IX−03/X/2007 (IBSP 174662); Cave N1_0015 (6°2'2"S, 50°16'16"W), 1♂7♀, 28/IX−03/X/2007 (IBSP 174665, IBSP 174664); Cave N1_0018 (6°2'1"S, 50°16'17"W), 2♂1♀, 28/IX−03/X/2007 (IBSP 174669) all collected by R. Andrade & I. Arnori et al.; Cave N1_0019 (6°2'1"S, 50°16'17"W), 1♀, 16/VII−06/VIII/2014, Equipe Carste et al. (IBSP 188834); Cave N1_0020 (6°1'53"S, 50°18'1"W), 1♀, 28/IX−03/X/2007 (IBSP 174674); Cave N1_0022 (6°1'57"S, 50°16'19"W), 2♂ 5♀, 28/IX−03/X/2007 (IBSP 174679, IBSP 174683); Cave N1_0025 (6°1'53"S, 50°16'20"W), 6♂17♀, 28/IX-03/X/2007 (IBSP 174685) all collected by R. Andrade & I. Arnori et al.; 1♂, 02-29/IV/2015, Equipe Carste et al. (IBSP 188847); Cave N1_0037 (6°1'50"S, 50°16'28"W), 2♀, 28/IX-03/X/2007, R. Andrade & I. Arnori et al. (IBSP 174688); Cave N1_0055 (6°1'12"S, 50°16'43"W), 1♀, 16/VII−06/VIII/2014, Equipe Carste et al. (IBSP 188831); Cave N1_0064 (6°1'7"S; 50°16'45"W), 3♀, 28/IX−03/X/2007 (IBSP 174692); Cave N1_0072 (6°1'13"S; 50°17'18"W), 4♀, 28/IX−03/X/2007 (IBSP 174698) all collected by R. Andrade & I. Arnori et al.; Cave N1_0073 (6°1'13"S; 50°17'17"W), 1♂5♀, 16/VII−06/VIII/2014 (IBSP 188845, IBSP 188833, IBSP 188838); 1♀, 02-29/IV/2015, Equipe Carste et al. (IBSP 188843); Cave N1_0075 (6°1'14"S, 50°16'49"W), 2♀, 28/IX-03/X/2007 (IBSP 174699, IBSP 174700); Cave N1_0098 (6°1'9"S, 50°17'5"W), 2♀, 28/IX-03/X/2007 (IBSP 174706); Cave N1_0103 (6°0'13"S, 50°17'55"W), 3♂10♀, 28/IX-03/X/2007 (IBSP 174707, IBSP 174708) all collected by R. Andrade & I. Arnori et al.; Cave N1_0109 (6°0'41"S, 50°18'41"W), 4♂2♀, 16/VII−06/VIII/2014, Equipe Carste et al. (IBSP 188848, IBSP 188846); Cave N1_0116 (6°0'39"S, 50°18'50"W) 1♂3♀, 28/IX−03/X/2007 (IBSP 174711), collected by R. Andrade & I. Arnori et al.; Cave N1_0141 (6°2'34"S, 50°16'32"W), 1♂10♀, 16/VII-06/VIII/2014 (IBSP 188840) collected by Equipe Carste et al.; Cave N1_0143 (6°1'36"S, 50°17'27"W), 3♂8♀, 28/IX−03/X/2007 (IBSP 174718, IBSP 174714); Cave N1_0170 (6°1'23"S, 50°17'58"W), 3♀, 28/IX−03/X/2007 (IBSP 174724); Cave N1_0173 (6°1'27"S, 50°17'55"W), 3♀, 28/IX−03/X/2007 (IBSP 174726) all collected by R. Andrade & I. Arnori et al.; Cave N1-0162 (6°0'55"S, 50°18'46"W), 1♀, 02-29/IV/2015 (IBSP 188841), collected by Equipe Carste et al.; Cave N2-026 (6°3'16"S, 50°14'23"W), 1♂1♀, 26/IX-17/X/2012 (IBSP 178499); 2♂3♀, 03-17/IV/2013 (IBSP 178502, IBSP 178501); Cave N3_0003 (6°1'44"S, 50°12'3"W), 1♂, 26/IX−17/X/2012 (IBSP 178481), 1♂1♀, 05-17/III/2013 (IBSP 178504, IBSP 178505); Cave N3_0004 (6°1'45"S, 50°12'2"W), 2♂0♀, 26/IX-17/X/2012 (IBSP 178482); Cave N3_0006 (6°1'45"S, 50°12'3"W), 1♂1♀, 26/IX−17/X/2012 (IBSP 178486, IBSP 178485) 1♂1♀, 05-17/III/2013 (IBSP 178506); Cave N3_0023 (6°2'35"S, 50°13'10"W), 4♂2♀, 05-17/III/2013 (IBSP 178508, IBSP 178507) 1♂1♀, 02-23/VIII/2013 (IBSP 178539); Cave N3_0026 (6°2'39"S, 50°13'9"W), 2♂, 26/IX−17/X/2012 (IBSP 178489, IBSP 178491); 4♂1♀, 05-17/III/2013 (IBSP 178509, IBSP 178512, IBSP 178510); Cave N3_0033 (6°2'42"S, 50°13'12"W), 2♂, 26/IX−17/X/2012 (IBSP 178492); 2♂3♀, 05-17/III/2013 (IBSP 178514, IBSP 178515); Cave N3_0036 (6°2'46"S, 50°13'13"W), 1♂0♀, 26/IX−17/X/2012 (IBSP 178494); Cave N3_0037 (6°2'45"S, 50°13'14"W), 1♀, 05-17/III/2013 (IBSP 178516); 1♂1♀, 26/IX−17/X/2012 (IBSP 178495); Cave N3_0039 (6°2'24"S, 50°13'21"W), 1♂0♀, 26/IX−17/X/2012 (IBSP 178497); Cave N3_0047 (6°2'27"S, 50°13'40"W), 3♂9♀, 03-17/IV/2013 (IBSP 178527, IBSP 178531, IBSP 178564, IBSP 178529, IBSP 178525, IBSP 178532); 2♂2♀, 02-23/VIII/2013 (IBSP 178541, IBSP 178540); Cave N3_0072 (6°2'36"S, 50°13'50"W), 2♀, 03-17/IV/2013 (IBSP 178533, IBSP 178534); Cave N3_0074 (6°2'35"S; 50°13'49"W), 6♂5♀, 05-17/III/2013 (IBSP 178518, IBSP 178517, IBSP 178521, IBSP 178519, IBSP 178524); 3♂, 02-23/VIII/2013 (IBSP 178546, IBSP 178548, IBSP 178549); Cave N3_0076 (6°2'28"S; 50°13'36"W), 1♂1♀, 03-17/IV/2013 (IBSP 178537); 2♂2♀, 02-23/VIII/2013 (IBSP 178552, IBSP 178553); all collected by Equipe Carste et al.; Cave N4E_0003 (6°2'25"S; 50°9'38"W), 3♀, 20/X-01/XI/2006 (IBSP 174942); Cave N4E_0007 (6°2'21"S; 50°9'36"W), 1♀, 20/X−01/XI/2006 (IBSP 174952); Cave N4E_0008 (6°2'21"S; 50°9'36"W), 9♀, 20/X−01/XI/2006 (IBSP 174954); Cave N4E_0010 (6°2'20"S; 50°9'38"W), 1♂9♀, 20/X−01/XI/2006 (IBSP 174968); 3♂5♀, 07-12/X/2008 (IBSP 174970, IBSP 174969); 4♂7♀, 20/IV-04/V/2010 (IBSP 176878, IBSP 176877, IBSP 176879, IBSP 176881, IBSP 176880); all collected by R. Andrade & I. Cizauskas et al.; Cave N4E_0011 (6°2'20"S; 50°9'38"W), 1♂6♀, 20/X−01/XI/2006, R. Andrade et al. (IBSP 174974); 3♀, 20/IV−04/V/2010 R. Andrade & I.Cizauskas et al. (IBSP 176883, IBSP 176882); Cave N4E_0012 (6°2'16"S; 50°9'37"W), 1♂0♀, 20/X−01/XI/2006 (IBSP 174976); Cave N4E_0013 (6°2'18"S; 50°9'38"W), 1♂7♀, 20/X−01/XI/2006 (IBSP 174979); all collected by R. Andrade et al.; 8♂16♀, 20/IV−04/V/2010 (IBSP 176886, IBSP 176888, IBSP 174981, IBSP 176885, IBSP 176884, IBSP 176889, IBSP 176887, IBSP 176890, IBSP 174060), collected by R. Andrade & I.Cizauskas et al.; Cave N4E_0014 (6°2'17"S; 50°9'37"W), 3♂4♀, 20/X−01/XI/2006 (IBSP 174987) 2♂6♀, 07-12/X/2008 (IBSP 174989, IBSP 174986, IBSP 174985); all collected by R. Andrade et al.; 3♂11♀, 20/IV−04/V/2010 (IBSP 176893, IBSP 176894, IBSP 176892, IBSP 176895, IBSP 176891); all collected by R. Andrade & I. Cizauskas et al.; Cave N4E_0015 (6°2'10"S; 50°9'35"W), 1♂3♀, 20/X−01/XI/2006 (IBSP 174991, IBSP 176876); Cave N4E_0016 (6°2'6"S; 50°9'37"W), 4♀, 20/X−01/XI/2006 (IBSP 174993); all collected by R. Andrade et al.; 1♂2♀, 20/IV−04/V/2010 (IBSP 176896, IBSP 176897); Cave N4E_0019 (6°2'4"S; 50°9'37"W), 2♀, 20/X-01/XI/2006 R. Andrade et al. (IBSP 175003); 3♂8♀, 20/IV−04/V/2010, R. Andrade & I. Cizauskas et al. (IBSP 176898, IBSP 176900, IBSP 176899); Cave N4E_0021 (6°2'2"S; 50°9'37"W), 6♂5♀, 20/X−01/XI/2006, R. Andrade et al. (IBSP 175010); 3♀, 20/IV-04/V/2010 (IBSP 176901, IBSP 176902, IBSP 176905); 3♂3♀, 20/IV−04/V/2010 (IBSP 176903, IBSP 176904); all collected by R. Andrade & I. Cizauskas et al.; Cave N4E_0022 (6°2'2"S; 50°10'4"W), 3♂7♀, 20/X−01/XI/2006 (IBSP 175012); 2♂3♀, 07-12/X/2008 (IBSP 175015); collected by R. Andrade et al.; 2♂7♀, 20/IV−04/V/2010 (IBSP 176906, IBSP 176908, IBSP 176907, IBSP 174061); all collected by R. Andrade & I. Cizauskas et al.; Cave N4E_0023 (6°2'1"S; 50°10'7"W), 1♂1♀, 20/IV−04/V/2010, R. Andrade & I. Cizauskas et al. (IBSP 176909); Cave N4E_0025 (6°2'1"S; 50°10'8"W), 1♀, 20/X−01/XI/2006, R. Andrade et al. IBSP 175024); Cave N4E_0026 (6°2'14"S; 50°10'3"W), 4♂10♀, 08-12/II/2007 R. Andrade & I. Arnori et al. (IBSP 175025); 4♂10♀, 08-12/II/2007 (IBSP 17502) 2♂4♀, 07-12/X/2008 R. Andrade et al. (IBSP 175028, IBSP 175033); 2♂7♀, 18/VIII-03/IX/2009, R. An drade & I. Cizauskas et al. (IBSP 176829, IBSP 176830, IBSP 176832, IBSP 176833, IBSP 176831); Cave N4E_0033 (6°2'25"S; 50°9'36"W) 4♂5♀, 08-12/II/2007, R. Andrade & I. Arnori et al. (IBSP 175060, IBSP 175053, IBSP 175055); 1♂7♀, 07-12/X/2008, R. Andrade & I. Cizauskas et al. (IBSP 175058, IBSP 175062, IBSP 175057); 2♂3♀, 18/VIII−03/IX/2009 (IBSP 176835) 8♂22♀, 15-22/IX/2009 (IBSP 176842, IBSP 176843, IBSP 176845, IBSP 176847, (IBSP 176849, IBSP 176836, IBSP 176846, IBSP 176844, IBSP 176848, IBSP 176837, IBSP 176841, IBSP 176838, IBSP 176839, IBSP 176840); all collected by R. Andrade & I. Cizauskas et al.; Cave N4E_0039 (6°1'58"S; 50°9'39"W) 2♀, 24-30/VII/2009 (IBSP 176850, IBSP 176851) 5♀, 19/II−04/III/2010 (IBSP 176866) collected by R. Andrade & I. Cizauskas et al.; Cave N4E_0041 (6°1'59"S; 50°9'42"W), 1♂1♀, 08-12/II/2007, R. Andrade & I. Arnori et al. (IBSP 175075); 1♂2♀, 24-30/VII/2009 R. Andrade & I. Cizauskas et al. (IBSP 176853, IBSP 176852); Cave N4E_0043 (6°1'55"S; 50°9'50"W) 1♀, 24-30/VII/2009 (IBSP 176854); Cave N4E_0044 (6°1'55"S; 50°9'50"W) 3♀, 24-30/VII/2009 (IBSP 176855, IBSP 176856) 3♀, 19/II−04/III/2010 (IBSP 176863, IBSP 176864); Cave N4E_0046 (6°2'16"S; 50°9'36"W) 1♂1♀, 19/II-04/III/2010 (IBSP 176868, IBSP 176867); Cave N4E_0051 (6°2'22"S; 50°9'38"W) 1♀, 24-30/VII/2009 (IBSP 176857); Cave N4E_0054 (6°2'1"S; 50°10'8"W) 1♂1♀, 19/II-04/III/2010 (IBSP 176865); all collected by R. Andrade & I. Cizauskas et al.; Cave N4E_0061 (6°2'21"S; 50°10'3"W) 4♂3♀, 08-12/II/2007, R. Andrade & I. Arnori et al. (IBSP 175080); 2♂9♀, 07-12/X/2008 (IBSP 175077, IBSP 175079); 2♀, 24-30/VII/2009 (IBSP 176861) 3♂7♀, 24-30/VII/2009 (IBSP 176858, IBSP 176860, IBSP 176859), all collected by R. Andrade et al.; Cave N4E_0062 (6°2'1"S; 50°9'12"W), 1♀, 24-30/VII/2009 (IBSP 176823); Cave N4E_0070 (6°1'56"S; 50°9'10"W), 1♀, 19/II−04/III/2010 (IBSP 176916); Cave N4E_0072 (6°1'56"S; 50°9'13"W), 6♀, 24-30/VII/2009 (IBSP 176826, IBSP 176827, IBSP 176828) 4♀, 19/II−04/III/2010 (IBSP 176874, IBSP 176875, IBSP 176872); Cave N4E_0080 (6°1'58"S; 50°9'4"W), 3♀, 19/II-04/III/2010 (IBSP 176869); Cave N4E_0089 (6°1'59"S; 50°9'6"W), 2♀, 24-30/VII/2009 (IBSP 176824, IBSP 176825); Cave N4WS-07 (6°5'22"S; 50°11'41"W), 1♂, 23/VIII/2010 (IBSP 176990); all collected by R. Andrade & I. Cizauskas et al.; Cave N4WS-08 (6°5'22"S; 50°11'41"W) 1♀, 07-12/X/2008, R. Andrade et al. (IBSP 174801); Cave N4WS-13 (6°3'59"S; 50°11'23"W), 2♂4♀, 20/X-01/XI/2006 (IBSP 174780); 1♂1♀, 20/IV-04/V/2010 (IBSP 176384, IBSP 176383); Cave N4WS-15 (6°3'59"S; 50°11'22"W), 14♂18♀, 20/X−01/XI/2006 (IBSP 174788, IBSP 174930, IBSP 174787, IBSP 174793, IBSP 174795, IBSP 174789), 3♂4♀, 07-12/X/2008 (IBSP 174812, IBSP 174808); all collected by R. Andrade et al.; 7♂24♀, 20/IV-04/V/2010 (IBSP 176386, IBSP 176394, IBSP 176395, IBSP 176396, IBSP 176398, IBSP 176399, IBSP 176388, IBSP 176389, IBSP 176393, IBSP 176392, IBSP 176387, IBSP 176391, IBSP 176385, IBSP 176397); all collected by R. Andrade & I. Cizauskas et al.; Cave N4WS-67 (6°4'22"S; 50°11'30"W), 1♂2♀, 18/XI−01/XII/2010 (IBSP 174070, IBSP 174069); all collected by C.R.A Souza & F.P. Franco et al.; Cave N5S-01 (6°5'27"S; 50°7'31"W), 1♀, 03/IV/2017, R. Zampaulo & X. Prous (IBSP 194763); Cave N5S-03 (6°6'18"S; 50°8'4"W) 1♂, 22/III−03/IV/2005, R. Andrade & I. Arnori et al. (IBSP 55367); 2♂1♀, 14-23/X/2009, R. Andrade & I. Cizauskas et al. (IBSP 177683, IBSP 177684, IBSP 177685); Cave N5S-05 (6°6'21"S; 50°8'1"W), 1♀, 22/III-03/IV/2005 (IBSP 55338); Cave N5S_06 (6°6'21"S; 50°8'2"W), 2♂1♀, 22/III−03/IV/2005 (IBSP 55344); Cave N5S-08 (6°6'21"S; 50°7'57"W), 1♂2♀, 22/III−03/IV/2005 (IBSP 55343); all collected by R. Andrade & I. Arnori et al.; 1♂, 07-12/X/2008 (IBSP 174524); 4♂5♀, 14-23/X/2009 (IBSP 177687, IBSP 177689, IBSP 177690, IBSP 177688); Cave N5S-09 (6°6'21"S; 50°7'53"W) 4♂12♀, 14-23/X/2009 (IBSP 177712, IBSP 177716, IBSP 177713, IBSP 177715, IBSP 177717, IBSP 177714); collected by R. Andrade & I. Cizauskas et al.; Cave N5S-10 (6°6'20"S; 50°7'53"W), 1♂, 22/III−03/IV/2005 (IBSP 55342), collected by R. Andrade & I. Arnori et al., 4♀, 07-12/X/2008 (IBSP 174526, IBSP 174528); 12♂23♀, 14-23/X/2009 (IBSP 177693, IBSP 177699, IBSP 177695, IBSP 177702, IBSP 177700, IBSP 177694, IBSP 177696, IBSP 177697, IBSP 177692, IBSP 177698, IBSP 177701, IBSP 177706); Cave N5S-11 (6°6'18"S; 50°7'47"W), 12♂25♀, 14-23/X/2009 (IBSP 177711, IBSP 177707, IBSP 177705, IBSP 177704, IBSP 177709, IBSP 177710, IBSP 177708, IBSP 177703); Cave N5S-14 (6°6'19"S; 50 °8'1"W), 1♀, 14-23/X/2009 (IBSP 177691); Cave N5S-15/16 (6°6'20"S; 50°7'60"W), 1♂, 14-23/X/2009 (IBSP 177686); Cave N5S-17 (6°5'15"S; 50°7'11"W), 1♂2♀, 25/VIII−03/IX/2009 (IBSP 177667, IBSP 177668, IBSP 177669); Cave N5S-18 (6°5'11"S; 50°7'39"W), 4♀, 25/VIII-03/IX/2009 (IBSP 177632, IBSP 177672, IBSP 177673, IBSP 177674); Cave N5S-19 (6°5'13"S; 50°7'37"W), 1♀, 25/VIII-03/IX/2009 (IBSP 177671); Cave N5S-20 (6°5'15"S; 50°7'35"W), 2♂7♀, 25/VIII-03/IX/2009 (IBSP 177651, IBSP 177652, IBSP 177653, IBSP 177654, IBSP 177655); Cave N5S-21 (6°5'15"S; 50°7'34"W), 2♂10♀, 07-12/X/2008 (IBSP 174533, IBSP 174537, IBSP 174541, IBSP 174534, IBSP 174536,IBSP 174538), 5♂11♀, 25/VIII-03/IX/2009 (IBSP 177656-IBSP 177658, IBSP 177660-IBSP 177666); Cave N5S-25 (6°5'12"S; 50°7'38"W) 1♂6♀, 14-16/XII/2010 (IBSP 188854, IBSP 178176); Cave N5S-30 (6°5'18"S; 50°7'10"W), 3♂8♀, 14-16/XII/2010 (IBSP 188855-IBSP 188859 4♂8♀, 10-19/V/2011 (IBSP 188860-IBSP 188867); collected by R. Andrade & I. Cizauskas et al.; Cave N5S-37 (6°6'22"S; 50°7'57"W), 1♂6♀, 07-12/X/2008, R. Andrade & et al. (IBSP 174543, IBSP 174545, IBSP 174542); 2♂20♀, 15-21/IX/2009 (IBSP 177679, IBSP 177718, IBSP 177722, IBSP 177681, IBSP 177723, IBSP 177680, IBSP 177678); 3♂4♀, 14/III-04/IV/2010 (IBSP 177721, IBSP 177719, IBSP 177720); Cave N5S-40 (6°6'19"S; 50°8'0"W), 1♀, 15-21/IX/2009 (IBSP 177724); Cave N5S-42 (6°6'21"S; 50°8'2"W), 1♂0♀, 25/VIII−03/IX/2009 (IBSP 177659); Cave N5S-52/53 (6°6'28"S; 50°7'59"W), 6♀, 25/VIII−03/IX/2009 (IBSP 177676, IBSP 177675, IBSP 177677); 1♂5♀, 14/III-04/IV/2010 (IBSP 177633, IBSP 177725, IBSP 177726, IBSP 177634); Cave N5S-57 (6°6'31"S; 50°7'57"W), 2♂6♀, 25/VIII−03/IX/2009 (IBSP 177635, IBSP 177636); 1♂3♀, 14/III−04/IV/2010 (IBSP 177727, IBSP 177728); Cave N5S-61 (6°6'19"S; 50°8'4"W), 1♀, 15-21/IX/2009 (IBSP 177729); Cave N5S-63/64/65 (6°6'12"S; 50°8'7"W) 1♀, 15-21/IX/2009 (IBSP 177682); 2♀, 14/III-04/IV/2010 (IBSP 177730, IBSP 177731); Cave N5S-84 (6°5'13"S; 50°8'12"W) 1♀, 25/VIII−03/IX/2009 (IBSP 177650); Cave N5S-85 (6°5'12"S; 50°7'35"W), 16♂20♀, 25/VIII−03/IX/2009 (IBSP 177637-IBSP 177649, IBSP 174066), 5♂7♀, 14/III−04/IV/2010 (IBSP 177732-IBSP 177735; IBSP 174065), all collected by R. Andrade & I. Cizauskas et al.; Cave N5SM1_0008 (-6,108887; -50,134387), 2♂5♀, 31/XIII/2010 (ISLA 14614) Cave N5SM1-05 (6°6'41"S; 50°8'7"W), 2♀, 01/IX/2010 (IBSP 176989); Cave N5SM1-13 (6°6'22"S; 50°8'6"W), 1♂2♀, 29/VIII/2010 (IBSP 176992); Cave N5SM1-21 (6°6'19"S; 50°8'16"W), 4♀, 02/IX/2010 (IBSP 176993); Cave N5SM1-38 (6°6'22"S; 50°8'12"W), 1♂3♀, 25/II/2011 (IBSP 176972); Cave N5SM1-42 (6°6'26"S; 50°8'6"W), 10♀, 19/II/2011 (IBSP 176999); Cave N5SM2_0003 (6°8'31"S; 50°8'6"W), 1♂1♀, 2010-11 (ISLA 14624); Cave N5SM2_0008 (6°8'27"S; 50°8'9"W), 1♀ (ISLA 14646); Cave N5SM2_0015 (6°8'17"S; 50°8'1"W), 1♂ (ISLA 14642); Cave N5SM2_0016 (6°8'17"S; 50°7'59"W), 1♂ (ISLA 14643); Cave N5SM2_0018 (6°8'18"S; 50°8'2"W), 2♂1♀ (ISLA 14638); Cave N5SM2_0022 (6°8'8"S; 50°8'7"W), 2♀ (ISLA 14627); Cave N5SM2_0023 (-6,135119; -50,13496), 1♀ (ISLA 14639); Cave N5SM2_0024 (6°8'8"S; 50°8'6"W), 1♂1♀ (ISLA 14644); Cave N5SM2_0025 (6°8'9"S; 50°8'6"W), 1♂, (ISLA 14632); Cave N5SM2_0026 (6°8'9"S; 50°8'6"W), 3♀ (ISLA 14621); Cave N5SM2_0037 (6°7'59"S; 50°8'5"W), 1♂3♀ (ISLA 14636); Cave N5SM2_0038 (6°7'58"S; 50°8'5"W), 1♂1♀ (ISLA 14647); Cave N5SM2_0042 (6°7'57"S; 50°8'11"W), 1♂5♀ (ISLA 14635, ISLA 14648); Cave N5SM2_0043 (6°7'56"S; 50°8'10"W), 1♂1♀ (ISLA 14628); Cave N5SM2_0044 (6°7'56"S; 50°8'6"W), 1♂1♀ (ISLA 14633); Cave N5SM2_0045 (6°7'55"S; 50°8'6"W), 1♂2♀ (ISLA 14641); Cave N5SM2_0053 (6°7'49"S; 50°8'5"W), 1♂1♀ (ISLA 14629); Cave N5SM2_0054 (6°7'48"S; 50°8'4"W), 1♀ (ISLA 14637); Cave N5SM2_0056 (6°7'47"S; 50°8'5"W), 3♀ (ISLA 14640); Cave N5SM2_0057 (6°7'47"S; 50°8'5"W), 1♂3♀ (ISLA 14620); Cave N5SM2_0058 (6°7'46"S; 50°8'5"W), 1♂4♀ (ISLA 14616); Cave N5SM2_0064 (6°7'43"S; 50°8'7"W), 1♀ (ISLA 14625); Cave N5SM2_0074 (6°7'32"S; 50°7'56"W), 2♂7♀ (ISLA 14623); Cave N5SM2_0076 (6°7'31"S; 50°7'54"W), 1♀ (ISLA 14645); Cave N5SM2_0078 (6°7'23"S; 50°7'49"W), 3♂5♀ (ISLA 14630, ISLA 14631); Cave N5SM2_0086 (6°7'16"S; 50°7'47"W), 1♂6♀ (ISLA 14618); Cave N5SM2_0087 (6°7'16"S; 50°7'43"W), 1♂1♀ (ISLA 14626); Cave N5SM2_0093 (6°7'17"S; 50°7'56"W), 6♂8♀ (ISLA 14619); Cave N5SM2_0096 (6°8'6"S; 50°8'12"W), 1♂1♀ (ISLA 14634), 2010-2011, all collected by Equipe UFLA; Cave N5W-03 (6°4'53"S; 50°8'4"W), 6♂2♀, 02-23/VIII/2013 (IBSP 178554, IBSP 178557, IBSP 178559, IBSP 178560, IBSP 178555, IBSP 178558); Cave N8-0024 (6°10'24"S; 50°9'8"W), 2♂5♀, 16/VII-06/VIII/2014 (IBSP 188830, IBSP 188832, IBSP 188835, IBSP 188839, IBSP 188844), 1♂, 02-29/IV/2015 (IBSP 188849); Cave N8-0026 (6°10'14"S; 50°9'28"W), 1♀, 16/VII−06/VIII/2014 (IBSP 188837), all collected by Equipe Carste et al.; Cave N5 E– 006 (6°05'05"S, 50°07'50"W), 1♂ 1♀, 22/ III– 03/V/2005, R. Andrade & I. Armoni (IBSP 55344); Cave N5E_0008 (6°04'54"S, 50°07'50"W), 1♂ 2♀, 22/ III– 03/V/2005, R. Andrade & I. Arnoni (IBSP 55343); Cave N5E_0005 (6°05'10"S, 50°07'48"W), 22/ III– 03/V/2005, 1♂ 1♀, R. Andrade & I. Arnoni (IBSP 55338); Cave N5E_0001 6°04'25"S, 50°07'05"W), 1♂, 22/ III– 03/V/2005, R. Andrade & I. Arnoni (IBSP 55342); EPIGEAN SAMPLES: Parauapebas, Floresta Nacional de Carajás, Serra Norte (6°5'19"S; 50°7'13"W), 2♀, 25/IV−03/V/2012 (IBSP 191339, IBSP 191357); (6°5'31"S; 50°7'35"W), 1♀, 25/IV−03/V/2012 (IBSP 191351); (6°6'12"S; 50°7'54"W), 7♂7♀, 27/I/2012 (IBSP 191340-IBSP 191348); (6°6'8"S; 50°7'54"W), 1♀, 26/IV−03/V/2012 (IBSP 191349); (6°6'17"S; 50°7'48"W), 1♂2♀, 26/IV−03/V/2012 (IBSP 191381, IBSP 191350), all collected by I. Cizauskas & R. Andrade et al.; (6°3'2"S; 50°14'56"W), 1♂, 10/X/2012 (IBSP 191393); (6°2'33"S; 50°13'6"W), 1♀, 28/IX/2012 (IBSP 191394); (6°3'9"S; 50°14'31"W), 1♀, 14/III/2013 (IBSP 191395), all collected by Equipe Carste et al.

Etymology.

The specific name refers to Varys, a fictional character in George R. R. Martin’s book, "A Song of Ice and Fire". Lorde Varys is a character with a venomous spirit, known as a spider in the plot.

Diagnosis.

Ochyrocera varys resembles Ochyrocera atlachnacha in its carapace, which is yellow and bright lime (Figs 1 A–B; 4 A–B) and palp with conical, elongated cymbial apophysis, and have a distal cuspule on the cymbial apophysis (Figs 1 C–D, 4 C–D). This species can be distinguished by the male palp having a cymbial apophysis without an accentuated lateral projection (present in O. atlachnacha ) and by the curved distal area of embolus (Figs 1 C–D, 2 C–F); females have a thick spermathecae enveloping large pore-plates (Fig. 1 E–F).

Description.

Male (holotype). Total length 2.3. Carapace length 0.7, ovoid; narrowing gradually anteriorly with yellow and bright lime pattern, fovea flattened and inconspicuous (Fig. 1A). Clypeus length 0.7. Eyes: PME oval; ALE and PLE rounded. Chelicerae light yellow; promargin with eight teeth, attached to long lamina (Fig. 2A); retromargin without teeth. Sternum yellow. Endites yellowish. Legs: light yellow; formula 1423; total length: I 7.0; II 5.9; III 4.1; IV 6.5. Male palp: palpal femur length 0.4; palpal tibia enlarged basally with several trichobothria; cymbial apophysis slightly curved distally, short apical cuspule, retrolateral paired long hair on non-projected base, a single tarsal organ, with two basal setae (Fig. 2 C–D), cymbial prolateral extension subtriangular (Fig. 2C); bulb oval; embolus elongated, wide at base and projecting upward, with sinuous tip (Fig. 2 C–F). Abdomen length 1.3, oval; uniformly grayish-green; six epiandrous spigots, with short base (Fig. 2B).

Female (paratype IBSP 176843). Total length: 2.0; carapace length: 0.74; Carapace pattern as in male (Fig. 1B). Pedipalp without claw, with conical tip and subdistal tarsal organ (Fig. 3 C–D). Clypeus: 0.67 diameter; Eyes, chelicerae, sternum, endites and labium as in male (Fig. 3B). Legs as in male; formula 4123, total length: I 6.3; II 4.7; III 3.6 IV4.3. Abdomen length 0.97. Colulus triangular with long bristles (Fig. 3A). Internal genitalia with well-developed medial columnar uterus externus, shorter than spermathecae length and internally with inconspicuous chambers. Uterus externus ending in a narrow neck. Rounded pore-plates covered by spermathecae, with approximately 15 glandular ducts (Fig. 1 E–F).

Distribution.

Recorded from caves and epigean areas of Carajás, state of Pará, northern Brazil (Fig. 19A).