Indopadilla vimedaba Maddison, sp. nov.

Maddison, Wayne P., Beattie, Imara, Marathe, Kiran, Ng, Paul Y. C., Kanesharatnam, Nilani, Benjamin, Suresh P. & Kunte, Krushnamegh, 2020, A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini), ZooKeys 1004, pp. 27-97 : 27

Indopadilla vimedaba Maddison, sp. nov. Figs 121-128

Stagetilus semiostrinus : Prószyński, 1987: figs on pages 105-105 (misidentification).

Padillothorax semiostrinus : Prószyński 2018: 174-175, figs 28F-L (misidentification).

Type material.

Holotype male (specimen JK 13.09.03.0011), in LKCNHM, from Singapore: Nee Soon Swamp, 2 September 2013. J. K. H. Koh.

Etymology.

An arbitrary combination of letters, ungendered. Other names: In Maddison (2015b) and WPM’s field or lab notebooks the informal code for this species was “BVMDB”.

Notes.

As noted under Padillothorax , this species was mistaken for Padillothorax semiostrinus , and illustrated by Prószyński (1987) under that name. He illustrates a male and female in the same vial 15151 in the MNHN Paris from the “Malayana” peninsula.

Diagnosis.

Palp with embolus longer that half the length of the bulb, straight and tapering (Fig. 121 ). Other Indopadilla have the embolus either longer and curved, or shorter. Epigyne with small but distinct ECP on back margin medially; openings simple and posterior (Fig. 123 ). Body dark brown except for highly contrasting but narrow pale cream markings, reticulate on abdomen.

Description.

Male (based on holotype, specimen JK 13.09.03.0011). Carapace length 2.4; abdomen length 3.5. Carapace in alcohol dark brown to black, slightly paler around fovea and in narrow medial band along thoracic slope, which also has some white scales; other patches of white scales along borders of ocular area. Clypeus extremely narrow at centre, and beneath it is a broad expanse of white arthrodial membrane; precise boundary between the clypeus and arthrodial membrane indistinct. Chelicerae with strong lateral ridge that extends into a flange near the fang, as drawn by Prószyński (1987: 105). Three promarginal teeth and at least five retromarginal. Palp honey coloured except darker base of femur. Embolus a narrow long triangle (Fig. 121 ). Legs honey coloured except much larger first leg, which bears a ventral fringe. Abdomen dark above and below, with two pair of lateral pale spots, the anterior of which connects with a few thin pale lines.

Female (based on specimen SWK12-3620). Carapace length 3.3; abdomen length 5.4. Carapace integument in alcohol medium red-brown except for narrow medial pale band along thoracic slope. Clypeus extremely narrow, exposing broad white arthrodial membrane (Fig. 128 ). Chelicerae concave in front with lateral ridge. At least four retromarginal teeth. Legs pale except first leg, dark except tarsus. Abdomen brown with distinct reticulation of pale scales. Epigyne (Fig. 123 ) with simple openings, from which broad copulatory ducts proceed anteriorly before narrowing considerably and proceeding to the posterior, then back anteriorly to the spermathecae.

Male-female matching.

The male and female described above were not co-collected, but they match well the male and female (MNHN 15151, photographs examined) described by Prószyński, which were in the same vial, and thus likely co-collected. These males and females match in markings: the first leg is all dark except the tarsus; the abdominal dorsum is dark with some narrow pale lines, the venter dark; the pale medial band of the integument of the thoracic slope is narrow; there is a narrow band of pale scales on the midline low on the thoracic slope; the triangle of scales near the fovea is narrow. The male has fewer lines in its abdominal markings than the female, but those it has are precise matches to the female. Other Indopadilla differ; e.g., another candidate female, I. nesinor , has the first leg considerably more strongly banded. suggesting her male should have a first patella paler than seen in Fig. 125 , more like that of I Indopadilla redunca ( Koh and Bay 2019:. 203, fig. E), whose female’s leg is banded similarly to I. nesinor . Also, I. nesinor differs in having a pale underside of the abdomen, and different thoracic markings. The relatively long but wide embolus of the male is a good fit to the openings and first broad part of the duct of the matched female, being ~ 0.07 mm wide at its base and ~ 0.3 mm long. In contrast, the openings of I. nesinor are only ~ 0.02 mm wide, enough to accommodate only a small portion of the embolus.

Additional material examined.

One female (specimen SWK12-3620, in UBCZ) from Malaysia: Sarawak: Mulu Nat. Pk., headquarters area, on & in buildings, 4.042°N, 114.814°E, 50 m el. 26-27 March 2012 Maddison/Ang WPM#12-100.

Figures

Figures 121 - 128. Indopadilla vimedaba sp. nov. 121 male left palp, ventral view (holotype JK. 13.09.03.0011) 122 same, retrolateral view of tibia 123 epigyne, ventral (specimen SWK 12 - 3620) 124 vulva, dorsal 125 holotype male 126 - 128 female SWK 12 - 3620. Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm.