Kamyristi yantardakhensis, Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024

Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent, 2024, Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities, Arthropod Systematics & amp; Phylogeny 82, pp. 151-181 : 151

3.2.1.1.2. † Kamyristi yantardakhensis sp. nov.

Figures 2H , 9D-F

Etymology.

The specific epithet refers to the locality where the specimen was collected. The specific adjective is to be treated as a noun in a genitive case.

Material studied.

Holotype male PIN 3311/2718; housed in the collection of the Paleontological Institute of the Russian Academy of Sciences (PIN).

Type locality.

Yantardakh, Russia.

Age.

Santonian, Upper Cretaceous.

Diagnosis.

Axillae not contiguous medially, grooves crenulated (Fig. 9F ; vs. contiguous medially, with axillar grooves smooth in † Kamyristi exfrigore gen. et sp. nov.).

Description.

Body length 1.47 mm. Body mostly bare. - Head slightly higher than long (length 0.26 mm; height 0.28 mm), glabrous; frons convex, shagreened; compound eye almond-shaped, higher than long, not covering head length, postocular carina absent; vertex convex, minutely shagreened but sculpture not fully visible; toruli inserted closer to each other than to eyes; subantennal groove present; mandible with three teeth, decreasing in size from apex to base; scape short and thick (length 0.07 mm), pedicel of similar length with scape but thinner; 10 flagellomeres preserved, cylindrical, elongate, longer than wide; flagellomere 1 shortest flagellomere (length 0.04 mm), flagellomeres 2-10 similar in length (length ca. 0.06 mm); flagellomere 11-12 missing; occipital carina crenulate. - Mesosoma more than a third of body length (length 0.43 mm; height 0.22 mm); mesoscutum less than one third of mesosoma length (length 0.15 mm; width 0.31 mm), shagreened and divided by crenulate median mesoscutal sulcus; axillae not continuous medially, with inner groove crenulate; mesoscutellum shagreened; pronotum not visible dorsally; anterior thoracic spiracle fully surrounded by pronotal cuticle; propodeum areolate-rugose. - Fore wing hyaline and covered with microtrichia (length ca. 0.75 mm); C, Sc+R, A, Rs and basal segment of M pigmented; M+Cu, Cu and 1m-cu spectral; Rs+M not aligned with M+Cu; medial cell pentagonal; Rs almost closing submarginal cell, not fully pigmented between r-rs and Rs+M; R1 pigmented beyond marginal cell; marginal cell closed by Rs with a bend; M pigmented halfway to wing tip and then spectral. - Legs bearing numerous scattered setae; one long mesotibial spur, one shorter metatibial spur; basimetatrochanter long, half metacoxa length; metafemur slightly swollen; first metatarsomere longest and bearing small erect setae. - Metasoma almost half body length, longer than mesosoma (length 0.75 mm), elongate; smooth and bare; tergites nearly equal in length; hypopygium small, parameres triangular, projecting downward.

Figures

Figure 2. Drawings of fore wing venation for the taxa described in the present paper. A † Cretolyra noijebumensis gen. et sp. nov.; B † Cretolyra shawi gen. et sp. nov.; C † Genkyhag innebula gen. et sp. nov.; D † Megacoxa chandrahrasa gen. et sp. nov.; E † Megacoxa janzeni gen. et sp. nov.; F † Megacoxa synchrotron gen. et sp. nov.; G † Kamyristi exfrigore gen. et sp. nov.; H † Kamyristi yantardakhensis gen. et sp. nov. Scale bars: 0.5 mm (A - H).

Figure 9. † Kamyristi gen. nov. A - C † Kamyristi exfrigore gen. et sp. nov., holotype female PIN 3730 / 411, Albian-Cenomanian, Taimyr amber. A habitus in lateral view; B habitus in dorsal view; C head and mesosoma in lateral view; D - F † Kamyristi yantardakhensis gen. et sp. nov., holotype male 3311 / 2718, Santonian, Taimyr amber; D habitus in lateral view; E habitus in ventral view; E habitus in dorsal view. Scale bars: 0.5 mm (A, B, D, E); 0.25 mm (C). AG = inner axillar groove; ce = cercus; MMS = median mesoscutal sulcus; OC = occipital carina; OvS = ovipositor sheaths; pa = parameres; sp = anterior thoracic spiracle.