Anchusa officinalis subsp. leucantha, 2013

Tan, Kit, Gjeta, Ermelinda, Mullaj, Alfred, Shuka, Lulezim & Vold, Gert, 2013, On the identity of Anchusa leucantha (Boraginaceae) from northern Greece, Phytotaxa 140 (1), pp. 35-42 : 37

Anchusa officinalis L. subsp. leucantha (Selvi & Bigazzi) Kit Tan , comb. et stat. nov. Figs. 1–5 .

Basionym: Anchusa leucantha Selvi & Bigazzi (2003: 436) . Type:― GREECE. Macedonia, nom. Edessa , colline rocciose c. 1 km a S di Drosia , suolo scistoso-ofiolitico, 2 May 2001, Bigazzi & Selvi 01.17 (holotype FI; isotypes ATH, B, G, W) .

Synonym : Anchusa officinalis var. ochroleuca Boissier (1875: 152) . Type :― F. Y. R. MACEDONIA. Prope Bitolia, 16 July 1862, Orphanides 136 (holotype G-Boiss.) .

Misapplied name: Anchusa ochroleuca sensu Grisebach (1844: 98) , non Bieberstein (1808: 125).

Plants caespitose with a rosette of basal leaves withered at anthesis. Basal leaves narrowly oblanceolate, obtuse or subacute, slightly undulate at margins. Cauline leaves linear to elliptic-lanceolate. Indumentum of simple spreading hairs up to 2.2 mm long, which are often tuberculate-bulbose at base on basal leaves. Cymes with single flower inserted at the branching points, branches elongating and lax in fruit. Calyx usually divided to half, distinctly pedicellate in fruit. Corolla creamy white, turning pale yellow when dry. Nutlets rugosetuberculate, reticulately-ridged and with thick basal ring.

Figures

FIGURE 1. Anchusa officinalis subsp. leucantha (holotype from eparchia Edessis, NC Greece).

FIGURE 2. Anchusa officinalis subsp. leucantha (from Florinis, NW Greece, determined by Selvi & Bigazzi as A. officinalis var. ochroleuca Boiss.).

FIGURE 3. Anchusa officinalis subsp. leucantha (from Librazhdi, SE Albania).

FIGURE 4. Anchusa officinalis subsp. leucantha (from Librazhdi, SE Albania): A, flowers; B, nutlets in lateral, ventral and dorsal view.

FIGURE 5. Distribution of Anchusa officinalis subsp. leucantha. The locus classicus of A. leucantha in eparchia Edessis is indicated by.