Canarium indistinctum Daly, Raharim. & Federman, 2015

Daly, Douglas C., Raharimampionona, Jeannie & Federman, Sarah, 2015, A revision of Canarium L. (Burseraceae) in Madagascar, Adansonia 37 (2), pp. 277-345 : 312-314

Canarium indistinctum Daly, Raharim. & Federman , sp. nov.

( Figs 6 ; 16 ).

Medium-sized trees, leaves 1-3-jugate, stipules 2-3 mm long, the stipular scar 1-1.5 mm long, lateral leaflet base usually symmetric, petiole and rachis with only sparse to scattered hairs; distinguished from C. elegans Daly, Raharim. & Federman , sp. nov. by the stipules 8-24 (vs 4-8 in C. elegans , sp. nov.) mm from petiole insertion, the lateral leaflets elliptic or less often oblong-elliptic or oblanceolate (vs ovate to slightly oblong-ovate), the fruit pedicel 4-10 (vs 3-4) mm long, and the fruit 3.2-5.2 × 1.7-2.6 cm (vs 2.8-3.5 × 1.7-1.9 cm) and (ob)ovoid to oblong-ellipsoid (vs (oblong-)obovoid and trigonous).

TYPUS. — Madagascar. Toamasina, path from “Pépinière” of Ambatondradama, Masoala Peninsula, Mahalevona , Maroantsetra , 375- 1011 m elev., 15°17’22”S, 50°00’58”E, 8.X.1997, C. Birkinshaw, H. Ralimanana & R. Ranaivojaona 494 (holo-, NY!; iso-, MO!, TAN). DISTRIBUTION AND ECOLOGY. — Known only from the type collection. So far as it is known, this species is a relatively small rainforest tree found at c. 950 m elevation near Maroantsetra on the Masoala Peninsula. Fruiting Jul-Oct .

ETYMOLOGY. — The specific epithet refers to the inconspicuous venation that characterizes this species.

DESCRIPTION

Trees to at least 15 m tall. Resin white when dry. Leaves 11- 21 cm long, 1-3-jugate; petiole 1.8-4.7 cm long, petiole and rachis with a few scattered flexuous to ascending white hairs to 0.3 mm long; stipules 4-8 mm from petiole insertion, c. 2 mm long, obovate and constricted at base, folded laterally, with dense appressed white hairs to 0.15 mm long, the scar 1-1.5 mm long and raised; basal petiolules 5-8 mm long, other laterals 4-12 mm long, the terminal one 12-30 mm long, petiolules deeply canaliculate, pulvinuli not conspicuous; basal leaflets 32-85 × 23-42 cm, broadly ovate to very broadly elliptic, other laterals 60-12.2 × 3.0- 4.4 cm, ovate to slightly oblong-ovate, terminal leaflet 6-10.4 × 3.2-4.6 cm, (slightly) (ob)ovate; leaflet apex abruptly and very broadly short-acuminate, the acumen 1-5 mm long; lateral leaflet base usually symmetric, sometimes slightly oblique, obtuse and slightly attenuate; margin flat; leaflets drying brown, coriaceous, dull; secondary vein framework festooned-brochidodromous, secondaries in 7-13 pairs, spreading or almost straight, spacing decreasing toward apex and base, the angle slightly increasing toward base, perpendicular epimedial tertiaries present (these reticulating distally), intercostal tertiaries random-reticulate and admedially branched, quaternaries random-reticulate; on abaxial side the midvein broadly prominent, rest of veins narrowly prominulous, with scattered appressed, thick, white hairs to 0.5 mm long on midvein and secondaries, plus a few on rest of surface; on adaxial surface the midvein narrowly prominulous but sunk in a groove, secondaries narrowly prominulous or flat, rest usually flat, sometimes broadly prominulous, surface glabrous except a few scattered ascending white hairs at base. Infructescence 5.5-11 cm long × 2.5 mm diam above base, axes with dense, flexuous, fine pale hairs to 0.15 mm long and sparse capitate glands; fruiting pedicel 3.7-4.2 × 3.7-4.2 mm, clavate, fruiting calyx 4-6 mm long, continuous with the pedicel, calyx lobes distinct, 2-3 mm long, rounded-triangular, nearly patent. Fruit 2.8-3.5 × 1.7- 1.9 cm [c. 4 × 2.5 cm when fresh], (oblong-)obovoid and trigonous, the apex rounded to slightly acuminate, the base acute to rounded; surface with dense, small, slightly raised, ferrugineous lenticels, also provided with descending, flexuous, white hairs to 0.4 mm long among lenticels, glabrescent.

NOTES

Canarium indistinctum , sp.nov. is distinguished from C. elegans , sp. nov. in the discussion under the latter species.

Figures

FIG. 6. — Distributions of Canarium betamponae Daly, Raharim. & Federman, sp. nov., C. bullatum (Leenh.) Daly, Raharim. & Federman, comb. et stat. nov., C. egregium Daly, Raharim. & Federman, sp. nov., and C. indistinctum Daly, Raharim. & Federman, sp. nov.

FIG. 16. — Canarium indistinctum Daly, Raharim. & Federman, sp. nov.: A, Fruiting branchlet with detail of fruit surface (A’), leaflet margin (A’’), and leaflet apex (A’’’); B, petiole base with stipular scars; A, B, Birkinshaw et al. 494 (NY). Scale bars: A, 2 cm; A’-A’’’, B, 5 mm.