Corymbitodes kambaitiana Fleutiaux, 1942

Schimmel, Rainer & Tarnawski, Dariusz, 2015, Three new species of the genus Corymbitodes Buysson, 1904 from Nepal and India with a description of the male of Corymbitodes kambaitiana Fleutiaux, 1942 (Coleoptera: Elateridae), Zootaxa 3925 (4), pp. 551-561 : 556-558

Corymbitodes kambaitiana Fleutiaux, 1942

(Plate III: figs 15–21a)

Type locality. Myanmar: Kambaiti.

Type material. Lectotype ♀: N.E. Burma ( Myanmar), 7000 ft., 3.IV.1934, R. Malaise; Ctenicera kambaitiana, Fleutiaux , type; subgenus Corymbitodes Buyss. , Metactenicerus Miwa , Syntypus; Lectotypus ♀, Corymbitodes kambaitiana Fleutiaux , des. Schimmel, 2014; Paralectotypes 5 ♀♀: same data as lectotype, 1 ♀, R. Malaise; same data but 12.IV.1934, 1 ♀, R. Malaise; same data but 8.IV.1934, 1 ♀, R. Malaise; same data but 19.IV.1934, 2 ♀♀, R. Malaise.

New material. Nepal: Ganesh Himal, 7 km W Godlang, 8.V 1.995,2950 m, 85o14’E, 28o10’N, 1 ♂, 1 ♀ ( CSV, HNHM), leg. G. Fábián & L. Ronkay.

Diagnosis. C. kambaitiana Fleutiaux, 1942 is closely related to C. chassaini sp. nov., but can easily be distinguished by the longitudinally campaniform pronotum, less dense pronotal punctures with interstices once to twice of puncture diameter, cuspidate pronotal hind angles, costiform arched interstices of elytral punctures, blackish sheen of the body and by the longer antennae, exceeding pronotal apices by the length of the last two antennomeres.

Description. ♂: Silky (dorsal and ventral sides), subparallel, slightly arched, with micro-reticulated integument, and eleven-segmented, moniliform antennae; pronotum longitudinally campaniform; elytra subparallel; length: 12.5 mm, width: 3.2 mm; black with blackish metallic sheen, legs and antennae blackish brown; pubescence very short, barely visible (fig. 15).

Head declined from base to apex, occiput with a short triangular impression; frons slightly elevated above base of antennae, apically edged; cranium with dense and umbilicate punctures, interstices of punctures reduced to small and arched wrinkles, some parts without punctures and distinctly matt; pubescence extremely short, almost invisible and declined to apex; eyes spherical; last segment of maxilla dolabriform; mandible bi-spicate, falciform apically; antennae moniliform from third antennomere on, exceeding pronotal apices by the length of the last two antennomeres; second antennomere subglobular, 1.5 times longer than wide apically, third antennomere subtrapezoidal, 2.2 times longer than wide apically and slightly longer than fourth and following antennomeres, antennomeres 4–10 subtrapezoidal, conspicuously obtuse and of decreasing apical width, all of them more than 1.8 times longer than wide apically (fig. 19), last antennomere oblong-elliptic, subapically very slightly constricted; pubescence short and bristly, especially long at apical edges.

Pronotum longitudinally campaniform, almost straight laterally, along median line distinctly longer than wide across posterior angles (length: 4.0 mm, width: 2.9 mm), slightly arched medially and declined to base; pronotum with a distinct oval median groove from base to apex, lateral edges very distinctly arched; posterior angles divergent, without carina (fig. 17), apically cuspidate (arrow, fig. 17); pronotal lateral edges completely visible in dorsal view; pronotal punctures medially and laterally umbilicate, medially with interstices reduced to small wrinkles, laterally with irregular interstices once to twice puncture diameter (fig. 16), pronotal surface microreticulated; pubescence bent from median line to apex and to lateral sides.

PLATE III. FIGURES 15–21. C. kambaitiana Fleutiaux, 1942 . 15—habitus, 16—pronotal punctures, 17—pronotal posterior angle, 18—elytral structure, 19—antennomeres 1–6, 20—elytral apex, 21—aedeagus, 21a—bursa copulatrix (paralectotype from Myanmar).

Pro-, meso- and metathorax finely punctate, with interstices plain and shiny; pubescence conspicuously long and accumbent.

Sides of mesosternal fossa gradually concave to body axis, declivous throughout length.

Prosternal apophysis straight, subparallel to body axis, with some transverse wrinkles, apex smoothly steplike, apically obtuse.

Sternites finely punctate and covered with bristly and accumbent hairs.

Scutellum lingulate, basally straight, apically arcuate, surface distinctly arched; punctures dense and rough, interstices reduced to small and arched wrinkles; pubescence dense, fine, directed from base to apex and to lateral sides.

Elytra subparallel, becoming very slightly narrower subapically, elytral apex arcuate (fig. 20), apically blunt without thorn (arrow, fig. 20); base very slightly shorter than pronotal posterior angles, slightly depressed at scutellum, margins arched, shoulders prominent (winged species); elytral striae with distinctly deep punctures, their interstices conspicuously costiform without punctures, very finely micro-reticulated and with distinct sheen (fig. 18); pubescence short, bristly, declined to apex and to lateral sides.

Alae transparent and densely covered with very small, almost invisible brownish bristles.

Legs elongate, moderately long and thin; trochanter trapezoidal, distinctly separated from femur; longitudinally oval femur with a distal groove for accommodation of tibia, with fine and short pubescence; tibiae covered with longer and protruding bristles; tarsomeres up to claws of decreasing length, ventrally with hardly visible, fine pubescence.

Mesosternal coxae proximally convex, gradually tapering to lateral sides.

Aedeagus long and elongate (fig. 21); penis subparallel, subapically distinctly constricted, slightly exceeding paramere apices; parameres apically conspicuously cochleariform, laterally slightly concave (arrow, fig. 21a).

Morphology of larvae unknown.

Etymology. Fleutiaux (1942) named this species for its type locality.

Distribution. Myanmar: Kambaiti; Nepal: Ganesh Himal.

Remarks. The species can be included in Group 2 mentioned in the discussion. The occurrence of C. kambaitiana in Nepal may appear enigmatic, as the type locality of the species in Myanmar (Kambaiti) is geographically distinctly separated from Ganesh Himal in Nepal, with geological dispersal barriers in between. However, the characters of the specimens from Nepal and the type specimens point to an absolute taxonomic congruence. Especially, the structure of the elytra with the costiform arched interstices of elytral striae punctures, and with very fine micro-reticulate distinctly sheen, are unique in the whole genus. This character is equally developed in males and females. Also, the reproductive organs are identically developed in the female specimens from Myanmar and from Nepal. The bursa copulatrix consists of an L-shaped, yellowish-transparent tube (length: 2.6 mm; width: 0.2–0.5 mm) covered with very fine, hardly visible sclerotised reddish-brown thorns (fig 21a); centrally on the tube there is a single structure of 5 parallel, digitiform scars (length: 0.1 mm).