Glaresis ecostata Fall, 1907

Gordon, Robert D. & Hanley, Guy A., 2014, Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea), Insecta Mundi 2014 (333), pp. 1-91 : 8-9

Glaresis ecostata Fall

Glaresis ecostata Fall 1907: 23 ; Gordon 1970: 502.

Description. Male. Length 3.2 mm, width 1.9 mm; body form elongate, slightly widened from elytral base to apical 1/3 ( Fig. 2A ). Color yellowish brown. Head with clypeus and frons smooth, shiny, finely micropunctate between large, deep punctures, large punctures separated by less than a diameter, setae short, barely emergent from punctures; frons densely microreticulate, dull, with small, round, widely spaced tubercles, apparently not setose; vertex with short, median basal carina, surface densely microreticulate, dull, without tubercles or punctures. Clypeal apex truncate, smooth, wide, angled posteriorly, posterior margin of angled apex raised, thickened ( Fig. 2B ), lateral angles feebly toothed. Mandible pair symmetrical; mesal tooth strong; lateral prominence strong, pronounced; outer margin angular. Pronotum with feeble central furrow extended from base nearly to apex, surface dull, finely, densely micoreticulate, with short, vertical, widely spaced seta-bearing tubercles, setae short, barely noticeable ( Fig. 2C ). Anterior and lateral pronotal margins not bordered, anterior margin smooth medially, crenulate in lateral 1/4, lateral and basal margins entirely crenulate. Elytron with flat striae lacking carinae, surface feebly shiny, distinctly, densely microreticulate; each stria with small, widely spaced, nearly round, seta-bearing tubercles, setae short, slightly shorter than diameter of an interval puncture; interval with large, round punctures somewhat obscured by microreticulate surface ( Fig. 2E ). Metasternum long, feebly shiny, finely, densely microreticulate, surface not tuberculate medially, laterally with some very long, oblique, seta-bearing tubercles (ridges); without trace of metasternal groove. Lateral protibial teeth unevenly spaced, basal tooth reduced. Mesotibia with 5 spines laterally, spinal bases close together, outer apex of tibia not strongly projecting ( Fig. 2D ). Posterior margin of metatrochanter with 2 large teeth, 1 small tooth ( Fig. 2H ); posterosuperior surface without teeth. Metafemoral surface with widely scattered, elongate, setae-bearing tubercles, finely microreticulate, shiny in apical 2/3, posterior 1/3 densely microreticulate, dull, slightly elongate; width to length ratio 1.0:1.4, without flange on anterior margin; posterosuperior margin with single large tooth, sometimes with additional small tooth. Metatibial surface finely microreticulate except extreme apex and apex of lateral expansion shiny (probably from wear); lateral metatibial margin serrate with row of small, seta-bearing teeth, posteromedian lateral projection large, shelf-like, apical margin uneven; row of more or less 3 tubercles anterior to lateral expansion; median surface and apex without tubercles; inner margin with row of small, seta-bearing tubercles, emarginate in apical 1/3, small tooth present at base of emargination ( Fig. 2G ). Apex of 5th adominal ventrite truncate. Genitalia long, dorso-ventrally flattened; basal piece shorter than parameres, proximal end curved; parameres straight in lateral view; median lobe much longer than parameres, slightly wider at middle than a paramere, strongly curved upward in apical 1/3, apex narrowly rounded in ventral and lateral views ( Fig. 2F ).

Female. Apex of 5th abdominal ventrite same as in male.

Variation. Length 3.2 to 5. mm, width 1.9 to 2.6 mm. Angled clypeal apex varies from narrow to wide; mesotibia with 5 or 6 lateral spines.

Type locality. California, Pilot Knob.

Type depository. MCZ.

Temporal distribution. April-June, September-October.

Geographical distribution. MEXICO. Baja California Norte: Campo Jemenez; Baja California Sur: San Felipe. Sonora: 6 mi N. El Golfo; vicinity of La Salina Ba. San Jorge, coastal sand dunes; Bahia Cholla; 6 mi. N. El Golfo. UNITED STATES. Arizona: Coconino Co., 1 mi. S Moenkopi; La Paz Co., Parker; La Paz Co., Cactus Plain Wilderness; La Paz Co., Route 95, 14 mi. SSE Parker; Mohave Co., Beaverdam; Navajo, 4 mi. N. Chilchinbito; Navajo Co., 1 mi. N Kayenta; Yuma Co., Ehrenberg; Co., San Luis; Yuma Co., Mohawk dunes. California: Imperial Co., 3 mi. N Glamis; 23 mi. W Winterhaven; Inyo Co., Saline Valley dunes; Algodones Dunes, 9.5 mi. NW Glamis; Algodones Dunes, Glamis, Osbourne Overlook; Holtville; Carrizo; Kern Co., Boron; Mojave Co., Beaverdam; Riverside Co., Chuckwalla Valley Dunes; Riverside Co., 2.8 km S. Hidden Palms; Riverside Co., Mecca; Riverside Co., Hopkins Well; Riverside Co., Indio; LaQinta; Palm Desert; San Bernardino Co., SE Amboy; San Bernardino Co., Amboy lava flow; Riverside Co., 18 mi. W Blythe; Riverside Co., Thousand Palms; San Bernardino Co., Baker; Death Valley, 4.3 mi. NE Saratoga; Kelso Dunes; 0 mi. E. 29 Palms; San Diego Co., Borrego; 1.1 mi. S Henderson Canyon Rd. on on Pegleg Rd.. Nevada: Nye Co., sand dunes in Amargosa Desert; Big Dune; Pershing Co., Woolsey. Utah: Washington Co., 2 mi. E Washington. ( BYUC) ( CASC) ( CICC) ( CMNC) ( CMNH) ( CNIC) ( CSCA) ( FSCA) ( KUNHM) ( MJPC) ( RAAC) ( RCCC) ( SMCC) ( TAMU) ( UCRC) ( USNM) ( WSCW).

Remarks. Glaresis ecostata is a large, smooth Glaresis readily distinguished from all other species by the key characters.

This species occurs primarily in Arizona and neighboring California, with its range extending south into Mexico and north to Nye Co., Nevada, and southern Utah. Its little sister species, G. costata , has a more extensive allopatric distribution to the east and north.

Figures

Figures 2A-2H. Glaresis ecostata Fall. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface. D) Metasternum and mesotibia. E) Elytral striae. F) Male genitalia. G) Metaleg and trochanter, dorsal view. H) Metaleg and trochanter, ventral view.