Glaresis gordoni Warner, 1995

Gordon, Robert D. & Hanley, Guy A., 2014, Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea), Insecta Mundi 2014 (333), pp. 1-91 : 41-42

Glaresis gordoni Warner

Glaresis gordoni Warner 1995: 268 ; Gordon 1970: 505 (as G. mendica ).

Description. Male. Length 2.7, width 1.4 mm; body form short, robust, slightly widened from elytral base to apical 1/3 ( Fig. 27A ). Color yellowish brown. Head with clypeal surface and frons densely rugose, reticulate with small, irregular ridges, with scattered large, irregular tubercles, setae short, indistinct; vertex without basal carina, surface rugose, reticulate with small, very irregular ridges, without tubercles or setae. Clypeal apex weakly emarginate, with small, evenly spaced tubercles, appearing dentate, lateral angles oblique, acute ( Fig. 27B ). Mandible pair symmetrical; mesal tooth strong; lateral prominence strong, pronounced; outer margin abruptly rounded. Pronotum with deep, long, transverse fovea in anterior 1/4 extended completely across pronotum, small, slightly transverse fovea on each side of middle in anterior 1/2, wide, deep fovea on each side medially near lateral margin, deep central furrow extended from base to transverse furrow; surface densely rugose, with sparse setae-bearing carinae, carinae mostly slightly irregular throughout, setae decumbent, about 1/2 length of carina ( Fig. 27C ); anterior and lateral pronotal margins not bordered, all margins crenulate. Elytra with surface feebly shiny, densely microreticulate; all striae with low carinae, carinae appearing continuous but narrowly interrupted adjacent to space between interval punctures, each carinal segment bearing an apical seta about 2/3 length of segment; intervals with deep round punctures. Metasternum long, feebly shiny, finely, densely microreticulate, median surface not tuberculate medially, laterally with irregular ridges in basal 1/2, median area slightly depressed, with feeble median carina extended anteriorly from apical keel nearly to middle; metasternal groove strong, deep, both sides abruptly ridged ( Fig. 27D ). Lateral protibial teeth unevenly spaced, basal teeth close together. Mesotibia with 3 short, widely spaced spines laterally, tibia slightly projecting at apex. Posterior metatrochanteral margin with 3 small teeth; posterosuperior surface of metatrochanter with 1 tooth, tooth not visible directly in ventral view ( Fig. 27G ). Metafemoral surface with widely scattered, elongate, setae-bearing tubercles, microreticulate; width to length ratio 1.0:1.5, with wide flange on anterior margin; posterosuperior margin without teeth or with faint suggestion of single tooth ( Fig. 27G ). Metatibia broadly triangular, surface entirely microreticulate, with posteromedian lateral projection, with row of small teeth on outer margin from near base to lateral projection, medially with irregular row of coarse tubercles extended from base nearly to projection, inner margin smooth, pubescent ( Fig. 27F ). Apical margin of 5th abdominal ventrite broadly rounded. Genitalia long, basal piece shorter than parameres, proximal end curved; median lobe shorter than parameres, as wide at middle as a paramere, slightly “pinched” medially, curved upward in apical 1/3, apex rounded in ventral view; parameres weakly curved in lateral view, slightly curved on inner margin, apex bluntly rounded ( Fig. 27E ).

Female. Apex of 5th abdominal ventrite same as in male.

Variation. Length 2.7 to 3.1 mm, width 1.4 to 1.6 mm. Head tubercles vary from small to pronounced; metasternal sculpture usually confined to basal 1/2 but occasionally present from base to apical 2/3; lateral margin of mesotibia occasionally with 4 teeth; posterosuperior surface of metafemur occasionally with small, indistinct tooth.

Type locality. Arizona, Yavapai Co., wash at NW side of Lake Pleasant.

Type depository. FSCA.

Temporal distribution. June-October.

Geographical distribution. MEXICO. Baja California Norte: 5.4 mi. NW Catavina; 49 mi. S Catavina; 12 km. NE El Arco; El Berrendo palm oasis; 12.7 km. east El Rosario; 14.7 km. east El Rosario; 30-35 mi. SE of El Rosario; 2.1 mi. SW Rosarito; La Zapopita; 9 km. NW rancho Santa Ines; 5 km. N. Ruben Jaramillo. Baja California Sur: 10 mi. N Cabo San Lucas; 34.6 mi. SE Guerrero; 8. mi. N Loreto; N. side of Loreto; 3.3 mi. N El Cien; 33.8 mi. N Loreto; 3 km. S. Los Barriles; 23 km. W La Paz; 8 mi. S. Mulege; 20 km. N Mulege, N. Palo Verde; 2 km. N El Progreso; 7 km. N Ranch Tablon. Sonora: 8 mi. S. Santa Ana; 2.5 mi. N. Hermosillo. UNITED STATES. Arizona: Cochise Co, 5 mi. N Benson; Cochise Co., San Simon Rd, 3.mi. N Portal; Tombstone; 5.8 km. SE Willcox, Blue Sky Road; Coconino Co., Sedona, Red Rock Crossing; Graham Co., Pinaleno Mts.; Maricopa Co., Jct Airport Rd. & Gila R., S. of Liberty; 10 mi. S Buckeye; Fort McDowell; Liberty; Tempe; Wickenberg; Pima Co., Baboquivari Mts.; Continental; Organ Pipe Cactus National Monument; Palo Verde Camp; Sabino Canyon, Santa Catalina Mts. ; Tucson Mts, Gilbert Ray Cmpg.; Pinal Co., Magma; Pinal Co., Superior; Santa Cruz, Co., Santa Rita Mts.; Yavapai Co., Camp Verde, Clear Creek Cmpg.; wash at NW side of Lake Pleasant. California: Riverside Co., Blythe; Cathedral City; Indio. Nevada: Clark Co., 15 mi. W. Searchlight; White’s City. New Mexico: Eddy Co., Carlsbad; San Juan Co., Cuaco Canyon. Utah: Washington Co., Zion National Park, Oak Creek. ( BYUC) ( CASC) ( CSCA) ( CICC) ( CMNC) ( CMNH) ( FSCA) ( JCIC) ( KUNHM) ( MJPC) ( SMCC) ( RAAC) ( RCCC) ( TAMU) ( UCRC) ( USNM) ( USUL).

Remarks. Glaresis gordoni is similar to G. tumida and G. caenulenta , but either completely lacks a tooth on the posterosuperior metafemoral margin, or occasionally with very small trace of a tooth, and has 3 small teeth on the hind trochanteral margin It is also unusual in this group because the lateral mesotibial margin nearly always has only 3 teeth.

Very frequently collected in southern Arizona, this species must also occur commonly south of the border. The Zion National Park site in Utah is distinctly disjunct from the primary known distribution of G. gordoni , as is the San Juan Co. locality in New Mexico, and these may be indicators of other seemingly isolated populations.

Figures

Figures 27A-27G. Glaresis gordoni Warner. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface. D) Metasternum with metasternal groove and mesotibia. E) Male genitalia. F) Metaleg and trochanter, dorsal view. G) Metaleg and trochanter, ventral view.