Magwengiella nycthemerops ( HEINRICH , 1978)

M, Alexander, 2015, A detailed illustrated description of Palearctic species Magwengiella (= Listrocalus) nycthemerops (HEINRICH, 1978). Notes on transformation of pigmental coloration of type specimens (Hymenoptera, Ichneumonidae, Ichneumoninae, Ctenocalini), Linzer biologische Beiträge 47 (2), pp. 1897-1907 : 1898-1900

Magwengiella nycthemerops ( HEINRICH, 1978) (Plate 1,2)

Listrocalus nyctemerops HEINRICH 1978 - East. Pal. Ichn.: 76-77,. Magwengiella nycthemerops BACHMAIER, DILLER 1985 – Entomofauna 6 (27): 490-491, comb.n. M a t e r i a l e x a m i n e d: Holotype:, Kazakhstan, Ural region ., Dzhanybek, 13.06.195 4,

RAFES P.M. leg. (MSU). Paratype:, Kazakhstan, lake. Chistoe at Semipalatinsk, 09.07.195 7,

PANFILOV D.V. leg. (ZSM).

M o r p h o l o g y

M a l e (holotype)

F l a g e l l u m: Short, bristle-shaped, with 26 segments, reddish-brown and without white annulus; segments (7)8-14 with weakly visible tyloides; segment of flagellum slightly ribbed ventrally. Flagellum 1,5 times shorter by length then front wing and 2 times shorter than body length.

H e a d: Head contour from front roundish, 1,1 times wider than height; length of genae (malar space) equal 0,3 of eye height; eyes not big; genae and temples visible up to upper third of an eye; head contour from above transversal, 2,1 times wider than length in middle and 1,8 times by an external contour. Vertex from lateral convex, roundly curved to occipital carina; temples long, 1,2 times longer than longitudinal diameter of an eye in the middle, temples from above swollen behind eyes, slightly and roundly narrowed backwards in hind half, from side parallel to hind margin of eye; occipital carina from above strongly roundly impressed but far not reach level of eyes and hind ocelli, sharp along the whole length, merged with hypostomal carina on the base of mandible; malar space 1,2 times longer than the mandible base width; mandible sickle-shaped, uniformly narrowed from base to apex, lower tooth moved inside and situated practically in same (horizontal) plane with upper one; clypeus transversal, almost two times wider than height, only just visible separated from middle field of face by very slight impression limiting middle field, convex at base and flattened to apex (slightly impressed) and lateral margins, with sharpened and distinctly broadly emarginated in middle front margin, lateral margins slanted, lateral angles rounded; labrum long, rounded, slightly narrower than front margin of clypeus with rather long pubescence; clypeal foveae small, rounded but distinct, area around them slightly impressed; surface of face slightly differentiated, but distinctly separated from genae, middle field of face more or less expressed, only just visible elevated in upper half above lateral fields and separated from lateral fields and clypeus by weak and very broad impressions; antennal cavities small, moderately and smoothly impressed, theirs upper borders reach level of front ocellus and lateral ones not reach borders of eyes, laterally above antennal fossae with distinct tubercles and without tubercle between antennal fossae; ocelli of normal size, diameter of lateral ocellus equal 0,6 distance to eye; ocellar triangle slightly elevated but more or less distinct. Face and base of clypeus punctured by small superficial punctures, surface between them polished, other part of head not sculpturated, smooth, polished, malar space very slightly shagreened.

T h o r a x: Collar of pronotum short with slightly rounded front margin, margin of collar above transverse furrow sharpened and high elevated, transverse furrow narrow and very deep, not interrupted by keel; margin of pronotal base slightly and uniformly curved; epomiae distinct but not sharp; pronotal ridge thickened, lateral impression far not reach upper margin. Mesonotum convex, 1,3 times longer than width; notauli distinct in front third; surface of mesonotum punctured by big smoothed irregular punctures (interspaces narrower than diameter of the punctures), shining, without microsculpture; axillary tongue not developed; prepectal carina sharp, at the top not reach front margin of mesopleurae; subalarum thick, high elevated; speculum impressed relative to front part of mesopleurae almost to level of area of mesopleural fovea, with sparce smoothed punctures, shining; area of mesopleural fovea deeply impressed relative to front part of mesopleurae; lower part of mesopleurae without distinct bend; sternauli absent; surface of mesopleurae densely punctured by smoothed punctures to smoothly wrinklypunctured at lover hind third, shining, without microsculpture; mesopleural suture interrupted by sharp ribs; scutellum high elevated, from behind vertically abrupt to postscutellum, horizontal part strongly convex with sparse punctures, laterally bordered by leaf-like broadened at horizontal part carinae; lateral foveae on postscutellum developed. Hind margin of metanotum with triangular projections opposite lateral longitudinal carinae of propodeum. Propodeum from lateral rather short, convex, without sharp bend between horizontal and vertical surfaces, length of horizontal part 2,2 times shorter than length of area posteromedia in middle; carinae of horizontal part of propodeum developed, with exception of lacking lateral carinae of basal area and only just marked external carina of area dentipara; lateral carinae of area posteromedia indistinct, coxal carina only just marked, practically absent; area superomedia transversal, 1,7 times wider than length in middle, almost horseshoe, costulae distinct, situated at middle of area superomedia; spiracles long, but not slit-shaped, length along external contour 2,7 times more than width in middle. Horizontal part of propodeum with big irregular punctures, except only just visible wrinkled area superomedia, shining, without microsculpture; area posteromedia sharply transversally-wrinkled; metapleurae with big irregular punctures, shining.

L e g s: Moderately short; coxae densely punctured by superficial smoothed punctures, shining; claws slightly broadened at base, curved in upper third.

W i n g s: Areolet quadrangular, asymmetrical, external veins of base some shorter than internal one and external veins of apex shorter than internal one; stigma rather narrow, light; radius practically straight; nervulus intersticial; ramulus rather long. Veins of both wings light, membrane of wings hyaline. Length of front wing 1,3 shorter than body length.

A b d o m e n:Fromabove broadly-oval, parallelsided up to apex of 4th tergite, tergites 2-4 strongly sclerotized with sharp constrictions between them, segments 6-7 visible from above; second tergite transversal, 1,4 times shorter than breadth at apex. First tergite from lateral with smoothed transition of petiolus to postpetiolus, dorsal and dorsolateral carinae of petiolus more or less distinct, merged at base of petiolus, ventrolateral carina expressed, lateral surface of petiolus with very weak ribs at ventrolateral carina; from above petiolus very narrow, sharply and roundly transform into broad postpetiolus; distance between spiracles 1,6 times more than distance from spiracles to hind margin of tergite; middle field of postpetiolus only just marked and not limited by carinae smoothly longitudinally wrinkly-punctured; lateral fields with sparse smoothed punctures, polished. Gastrocoeli short, deep, 1,3 times less by width than interval between them; thyridia indistinct, but expressed; interval with sharp longitudinal wrinkles extend almost behind middle of tergite; lunulae in a form of weak impressions behind middle of tergite; surface of second tergite except middle punctured by big smoothed punctures; tergite 3 from base to middle medially and 4th at base roughly wrinkly-punctured, laterally both tergites punctured by small smoothed punctures; tergites 5-7 smoothed and shining with very delicate punctures. Hypopygium short, triangle with rounded apex; all sternites of abdomen with longitudinal fold.

S i z e: Body length – 8,6; front wing – 6,6; flagellum – 4,3 mm.

C o l o r a t i o n

D i s c u s s i o n

As noted above, the description of the coloration of the holotype by the author of the original description does not correspond with coloration of sample with the geographical label of holotype. In our view, when describing the coloration of species, G. Heinrich (1978) used both the holotype and the paratype specimens. The description thus is of a compromise nature. Comparative table with description of the coloration of parts and segments of a body given by Gerd Heinrich (1978), description of the holotype coloration at present time and coloration of the paratype from Zoologische Staatssammlung, Munich done based on its photo made by Dr. Stefan Schmidt (in 2013) is given below (Table).